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Vincent Kalkman

Dr. Ing. Vincent J. Kalkman - Insect biodiversity


Email: vincent.kalkman@naturalis.nl
Phone: +31 (0)71-7517373
Room number: Darwinweg 2
“There are few countries that offer better possibilities for biodiversity studies than the Netherlands.”

The tropics, with its wealth of species, is often thought to be the best place to study biodiversity. Difficult field conditions, taxonomical problems and lack of basic data on for instance distribution or abiotic factors make work on biodiversity in the tropics however often difficult. The Netherlands is less species rich but has with 40.000 species still a bewildering diversity on offer. The amount of information available on the Dutch flora and fauna is almost unrivaled and information on land use, climate and human impact is extremely detailed. The possibilities offered by the Netherlands for studies on biodiversity are nearly endless but are still poorly explored.


I currently work as scientist for the EIS Kenniscentrum insecten and a program leader Bhutan for Naturalis Biodiversity Center

EIS Kenniscentrum Insecten brings together information on distribution and ecology of invertebrates.  In order to do so we maintain distribution databases, organize mapping schemes and publish a newsletter, a journal, a series of entomological keys and a book series. Much of the information gathered by the EIS can be found through the website Dutch Species Catalogue (Nederlands Soortenregister), a complete list of the biodiversity of the Netherlands. In addition we act as consultant for governmental and non-governmental organizations giving advice on policy and conservation regarding invertebrates in the Netherlands. This includes the production of red lists, conservation action plans and advice on invasive species. 

Naturalis Biodiversity Center, Leiden is actively involved in the study of Bhutanese biodiversity. This is done in cooperation with the Ugyen Wangchuck Institute for Conservation and Environment (UWICE), College of Natural Resources (CNR) at Lobesa, Sherubtse College and the National Plant Protection Centre and is coordinated by the Bhutanese National Biodiversity Center, Thimphu. The project is funded by the Bhutan Trust Fund for environment and conservation and the Uyttenboogaart-Eliasen Foundation. This cooperation has resulted in a series joined field expeditions held since 2013 focusing on wasp, dragonflies, moths, ladybeetles and mollusc. In the period 2017-2020 we hope to put more emphasis on freshwater invertebrates and on pollinators. 


Research interest

Biodiversity of the Netherlands & Himalayan region

Bumblebee conservation

Image recognition & biodiversity research

Taxonomy en phylogeny of Dragonfly & dragonfly of New Guinea


entomology, netherlands, biodiversity, odonata, freshwater, bhutan, new guinea

Current research topics






Available student projects


Naturalis Repository



Kalkman, V.J., J.-P. Boudot, R. Bernard, G. De Knijf, F. Suhling & T. Termaat 2018. Diversity and conservation of European dragonflies and damselflies (Odonata). Hydrobiologia 811: 269-282, doi.org/10.1007/s10750-017-3495-6. PDF

Kalkman, V.J. K-D. B. Dijkstra, R.A. Dow, F.R. Stokvis & J. van Tol 2017 Out of Australia: the Argiolestidae reveal the Melanesian Arc System and East Papua Composite Terrane as possible ancient dispersal routes to the IndoAustralian Archipelago (Odonata: Argiolestidae), International Journal of Odonatology 21: 1-14, DOI: 10.1080/13887890.2017.1402825



Cuppen, J.G.M., V.J. Kalkman & G. Tacoma-Krist 2017 Verspreiding, biotoop en fenologie van de Nederlandse lieveheersbeestjes (Coleoptera: Coccinellidae). Entomologische Berichten 77(3) 147-187.

Gyeltshen, T. & V.J. Kalkman 2017 Odonata records from western Bhutan, with six new records and a note on the synonymy of Himalagrion with Coenagrion. Notulae odonatologicae 8(9): 341-353.

Gyeltshen, T., T. Nidup, P. Dorji, T. Dorji & V.J. Kalkman 2017 A bibliography and checklist of the dragonflies and damselflies of BhutanJournal of Threatened Taxa 9: 9743–9747

Gyeltshen, T., V.J. Kalkman & A.G. Orr 2017 Honouring His Royal Highness the Crown Prince of Bhutan: Megalestes gyalsey (Odonata: Synlestidae). Zootaxa 4244: 588–594.

Gyeltshen, T., V.J. Kalkman & A.G. Orr 2017 Field Guide to the common dragonflies & damselflies of Bhutan. National Biodiversity Centre, Thimphu.



Kalkman, V.J. 2016 Revision of the genus Celebargiolestes Kennedy, 1925 (Odonata: Argiolestidae). Odonatologica 45: 235-269.

Kalkman, V.J. & A.G. Orr 2016 A description of the larva and discussion of radiation in the phytotelm breeding damselfly genus Papuagrion in New Guinea (Odonata: Zygoptera: Coenagrionidae). International Journal of Odonatology 19: 169-182 doi: 10.1080/13887890.2016.1203363

Kalkman, V.J. & T. Gyeltshen 2016 Records of dragonflies from western Bhutan collected in October 2015. IDF-report 94: 1-20

Orr, A.G. & V.J. Kalkman 2016 Two new species of Papuagrion Ris, 1913 (Odonata: Coenagrionidae) from New Guinea with a survey of distribution records for the genus. Zootaxa 4173: 18–28.

van Strien, A.J.,  A.W. Gmelig Meyling, J.E. Herder, H. Hollander, V.J. Kalkman, M.J.M. Poot, S. Turnhout, B. van der Hoorn, W.T.F.H. van Strien-van Liempth, C.A.M. van Swaay, C.A.M. van Turnhout, R.J.T. Verweij, N.J. Oerlemans 2016. Modest recovery of biodiversity in a western European country: The Living Planet Index for the Netherlands. Biological Conservation 200: 44–50. doi:10.1016/j.biocon.2016.05.031



Boudot, J.P. & V.J. Kalkman 2015 Atlas of the European dragonflies and damselflies. KNNV Uitgeverij, Utrecht.

Orr, A.G. & V.J. Kalkman 2015 Field Guide to the dragonflies of New Guinea. Brachytron 17: 1-154.

Orr, A.G. & V.J. Kalkman 2015 Nannophlebia leoboppi sp nov., a new dragonfly species from New Guinea (Odonata: Anisoptera: Libellulidae). Zootaxa 3964 (3): 391–395.

Suhling, F., G. Sahlén, S. Gorb, V.J. Kalkman, K-D.B. Dijkstra, J. van Tol 2015 Order Odonata. In: Thorp, J., Rogers, D.C. (Eds.), Ecology and General Biology: Thorp and Covich's Freshwater Invertebrates, Academic Press, 893–932. ISBN: 9780123850263



Boudot, J.-P. & V.J. Kalkman 2014 Chapter 5. Odonata (dragonflies and damselflies). In: Smith et al. The status and distribution of freshwater biodiversity in the Eastern Mediterranean. IUCN, Cambridge. 

Kalkman, V.J. & A.G. Orr 2014 Distribution and identification of Rhodothemis in the eastern part of the Indo-Australian Archipelago (Odonata: Libellulidae) Faunistic Studies in South-East Asian and Pacific Island Odonata 8: 1-9

Kalkman, V.J. & T. Termaat 2014 Wat vertellen de SNL-insecten ons? Dagvlinders, libellen en sprinkhanen geven een gemengd beeld. Vakblad natuur bos landschap 108: 10-13.

Theischinger, G. & V.J. Kalkman 2014 The genus Teinobasis on the Bird's Head Peninsula and the Raja Ampat Islands, Indonesia (Odonata: Coenagrionidae). Odonatologica 43: 143-168.

Theischinger, G. & V.J. Kalkman 2014 Nososticta rufipes sp. nov. from Misool, with notes on the other Nososticta species on the Bird's Head Peninsula and the Raja Ampat, Indonesia (Zygoptera: Platycnemididae). International Journal of Odonatology 17: 217-222



Dijkstra, K.-D.B., G. Bechly, S.M. Bybee, R.A. Dow, H.J. Dumont, G. Fleck, R.W. Garrison, M. Hämäläinen, V.J. Kalkman, H. Karube, M.L. May, A.G. Orr, D. Paulson, A.C. Rehn, G. Theischinger, J.W.H. Trueman, J. van Tol, N. von Ellenrieder & J. Ware 2013 The classification and diversity of dragonflies and damselflies (Odonata). In: Zhang, Z.-Q. (Editor). Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3703: 36-45.

Dijkstra, K.-D.B., & V.J. Kalkman 2013 The ‘African’ genus Argiagrion is a Brazilian Leptagrionspecies and the ‘Philippine’ Moroagrion a European Pyrrhosoma (Odonata: Coenagrionidae). International Journal of Odonatology 16: 189-191.

Dijkstra, K.-D.B., V.J. Kalkman, R.A. Dow, F.R. Stokvis & J. van Tol 2013 Redefining the damselfly families: a comprehensive molecular phylogeny of Zygoptera (Odonata). Systematic Entomology 39: 68–96 DOI: 10.1111/syen.12035

Kalkman, V.J. & A.G. Orr 2013 Field Guide to the damselflies of New Guinea. Brachytron 15: 3-120.

Kalkman, V.J. & G. Theischinger 2013 Generic revision of Argiolestidae (Odonata), with four new genera, International Journal of Odonatology 16: 1-52.

Kalkman, V.J., S.J. Richards & D.A. Polhemus 2013 Two new species of Pyrrhargiolestes, with a key to the males (Odonata: Argiolestidae), International Journal of Odonatology 16: 53-65.

Musters, K., V.J. Kalkman & A. van Strien 2013 Predicting rarity and trend of animals, plants and fungi using a universal list of traits and indicator groups. Ecology and Evolution: 3: 3401–3414 DOI:  10.1002/ece3.699

Kalkman, V.J., K.-D.B. Dijkstra, R.A. Dow, F.R. Stokvis & J. van Tol 2013 Out of Australia, aquatic insects reveal the Melanesian Arc System and East Papua Composite Terrane as pathway to the Malesian region (Odonata, Argiolestidae). PhD-thesis.

Orr, A.G., V.J. Kalkman & S.J. Richards 2014 Four new species of Palaiargia Förster, 1903 (Odonata: Platycnemididae) from New Guinea with revised distribution records for the genus. International Journal of Odonatology 16: 309-325.

Van Strien, A.J., T. Termaat, V.J. Kalkman, Prins, M. De Knijf, G. Gourmand A.-L., Houard, X. Nelson, B. Plate, C., Prentice, S. Regan, E., Smallshire, D. Vanappelghem, C., Vanreuse, W. 2013 Occupancy modelling as a new approach to assess supranational trends using opportunistic data: a pilot study for the damselfly Calopteryx splendens. Biodiversity Conservation 22: 673-686 



Bilz, M., A. Nieto, S. Sánchez, K.N.A. Alexander, A. Cuttelod, V.J. Kalkman, E. Neubert, M. Seddon, C. van Swaay (2012) Invertebrates: our natural capital. In: Collen B, Böhm M, Kemp R & Baillie J.E.M. 2012 Spineless: status and trends of the world’s invertebrates. Zoological Society of London, United Kingdom. PDF

Dijkstra, K.-D.B. & V.J. Kalkman 2012 Phylogeny, classification and taxonomy of European dragonflies and damselflies (Odonata): a review. Organisms Diversity & Evolution 12: 209-227, DOI 10.1007/s13127-012-0080-8.

Kalkman, V.J., R.M.J.C. Kleukers & J. Tiago Tavares 2012 First well documented records ofOrthetrum trinacria for Greece and Turkey (Odonata: Libellulidae). Libellula: 31: 89-96.

Kalkman, V.J. & A.G. Orr 2012 The Australian monsoon tropics as a barrier for exchange of dragonflies (Insecta: Odonata) between New Guinea and Australia. Hydrobiologia 693: 55-70. DOI 10.1007/s10750-012-1086-0.

Orr, A.G., V.J. Kalkman & S.J. Richards 2012 A review of the new Guinean genus Paramecocnemis Lieftinck (Odonata: Platycnemididae), with the description of three new species. Australian Entomologist 39: 161-177.

Termaat, T. & V.J. Kalkman 2012 Basisrapport Rode Lijst Libellen 2011 volgens Nederlandse en IUCN-criteria. Brachytron 14: 75-186.



De Knijf, G. U. Flenker, C. Vanappelghem, C.O. Manci, V.J. Kalkman & H. Demolder 2011 The status of two boreo-alpine species, Somatochlora alpestris and S. arctica, in Romania and their vulnerability to the impact of climate change Odonata: Corduliidae). International Journal of Odonatology 14: 111-126.

Kaize, J. & V.J. Kalkman 2011 Records of dragonflies (Odonata) from Kabupaten Asmat and Kabupaten Mappi (Papua, Indonesia). Suara Serangga Papua 2011 5: 99-107. 

Kalkman, V.J., G. Theischinger & S.J. Richards 2011 Dragonflies and damselflies of the Muller Range, Papua New Guinea. In: Richards. S.J. & B.G. Gamui (ed.) 2011. Rapid Biological Assessments of the Nakanai Mountains and the Upper Strickland Basin: surveying the biodiversity of Papua New Guinea's sublime karst enviroments. RAP Bulletin of Biological Assessment 60. Conservation International. Arlington, VA. PDF

Kalkman, V.J. & R.J.T. Villanueva 2011 A synopsis of the genus Rhinagrion with description of two new species from the Philippines (Odonata: Megapodagrionidae). International Journal of Odonatology 14: 11-31.   

Zhang, H., V.J. Kalkman & X. Tonga 2011 A synopsis of the genus Philosina with descriptions of the larvae of P. alba and P. buchi (Odonata: Megapodagrionidae).International Journal of Odonatology 14: 55 – 68.



Balian, E. I.J. Harrison, H. Barber-James, S.H.M. Butchart, P.Chambers, J. Cordeiro, N. Cumberlidge, F. de Moor, C. Gascon, V.J. Kalkman, P.P. van Dijk, D. Yeo. 2010 A wealth of life, species diversity in freshwater systems. In: Mittermeier, R.A., T.A. Farrell, I.J. Harrison, A.J. Upgen, T.M. Brooks. Freshwater, the essence of life, Cemex conservation book series.

Kaize, J. & V.J. Kalkman 2010 On a collection of dragonflies (Odonata) from the Island of Mioswaar (Papua Barat, Indonesia). Suara Serangga Papua 5: 71-76.

Kalkman, V.J., C.Y. Choong, A.G. Orr & K. Schütte 2010 Remarks on the taxonomy of Megapodagrionidae with emphasis on the larval gills (Odonata). International Journal of Odonatology 13: 119-135. 

Kalkman, V.J. & K.D.P. Wilson 2010. Calilestes and Lestomima, junior synonyms of Rhipidolestes (Odonata: Megapodagrionidae). International Journal of Odonatology 13: 97-102. 

Kalkman, V.J., S.J. Richards & D.A. Polhemus 2010. Three new species of Argiolestes, with a key to the males of Argiolestes s. str. (Odonata: Megapodagrionidae). International Journal of Odonatology 13: 75-88, pls II, IIIa. 

Kalkman, V.J., J.-P. Boudot, R. Bernard, K.-J. Conze, G. De Knijf, E. Dyatlova, S. Ferreira,M. Jović, J. Ott, E. Riservato and G. Sahlén. 2010. European Red List of Dragonflies. Luxembourg: Publications Office of the European Union. PDF

Kalkman, V.J., B. Odé & A.W. Gmelig Meyling 2010 Patronen in de Nederlandse biodiversiteit. – In: Noordijk, J., R.M.J.C. Kleukers, E.J. van Nieukerken & A.J. van Loon (redactie). De Nederlandse biodiversiteit. Nederlandse Fauna 10. Nederlands Centrum voor Biodiversiteit Naturalis & European Invertebrate Survey-Nederland, Leiden: 318-338.

Kalkman, V.J., L. van Duuren, A.W. Gmelig Meyling & B. Odé 2010 Veranderingen in de Nederlandse biodiversiteit. – In: Noordijk, J., R.M.J.C. Kleukers, E.J. van Nieukerken & A.J. van Loon (redactie). De Nederlandse biodiversiteit. Nederlandse Fauna 10. Nederlands Centrum voor Biodiversiteit Naturalis & European Invertebrate Survey-Nederland, Leiden: 339-354.

Kalkman, V.J. 2010 Odonata - libellen.  – In: Noordijk, J., R.M.J.C. Kleukers, E.J. van Nieukerken & A.J. van Loon (redactie). De Nederlandse biodiversiteit. Nederlandse Fauna 10. Nederlands Centrum voor Biodiversiteit Naturalis & European Invertebrate Survey-Nederland, Leiden: 203-205.

Orr, A.G. & Kalkman, V.J. 2010 Arrhenocnemis parvibullis sp. nov. (Odonata: Platycnemididae), a new calicnemiine damselfly from Papua New Guinea, with a description of the female of A. amphidactylis Lieftinck, 1949. The Australian Entomologist 37: 137-146. 

Schouten, M.A., A. Barendregt, P.A. Verweij, V.J. Kalkman, R.M.J.C. Kleukers, H.J.R. Lenders & H.N. Siebel 2010 Defining hotspots of characteristic species for mutiple taxonomic groups in the Netherlands. Biodiversity and conservation 19: 2517-2536. 

Samraoui, B., J.B. Boudot, S. Ferreira, E. Riservato, M. Jović, V.J. Kalkman & W. Schneider 2010 The status and distribution of dragonflies. In: García, N., Cuttelod, A. and Abdul Malak, D. (eds.) (2010). The Status and Distribution of Freshwater Biodiversity in Northern Africa. Gland, Switzerland, Cambridge, UK, and Malaga, Spain : IUCN, 2010. PDF

Termaat, T., V.J. Kalkman, J.H. Bouwman 2010 Changes in the range of dragonflies in the Netherlands and the possible role of temperature change. BioRisk 5: 155–173. 



Boudot, J.-P., V.J. Kalkman, M. Azpilicueta Amorín, T. Bogdanović, A. Cordero Rivera, G. Degabriele, J.-L. Dommanget, S. Ferreira, B. Garrigós, M. Jović, M. Kotarac, W. Lopau, M. Marinov, N. Mihoković, E. Riservato, B. Samraoui & W. Schneider 2009 Atlas of the Odonata of the Mediterranean and North Africa. Libellula Supplement  9: 1-256.

Clausnitzer, V., V.J. Kalkman, M. Ram, B. Collen, J.E.M. Baillie, M. Bedjanič, W.R.T. Darwall, K.-D.B. Dijkstra, R. Dow, J. Hawking, H. Karube, E. Malikova, D. Paulson, K. Schütte, F. Suhling, R. Villanueva, N. von Ellenrieder & K. Wilson 2009 Odonata enter the biodiversity crisis debate: the first global assessment of an insect group. Biological Conservation 142: 1864-1869. 

Kalkman, V.J., H. van Mastrigt & S.J. Richards 2009 First records of dragonflies (Odonata) from the Foja Mountains, Papua Province, Indonesia. Suara Serangga Papua 4 (1) 14-18.

Kaize, J. & V.J. Kalkman, 2009 Records of dragonflies from kabupaten Merauke, Papua, Indonesia collected in 2007 and 2008 (Odonata). Suara Serangga Papua, 4 (2) 40-45. 

Schouten, M.A., P. A. Verweij, A. Barendregt, R. M. J. C. Kleukers, V. J. Kalkman, P. C. de Ruiter 2009 Determinants of species richness patterns in the Netherlands across multiple taxonomic groups. Biodiversity and Conservation, 18: 203-217. 

Riservato, E., J.P. Boudot, S. Ferreira, M. Jović, V.J. Kalkman, W. Schneider, B. Samraoui & A. Cuttelod 2009 The Status and Distribution of Dragonflies of the Mediterranean Basin.Gland, Switzerland and Malaga, Spain: IUCN. vii + 33 pp. PDF



Bouwman, J.H., V.J. Kalkman, G. Abbingh, E.P. de Boer, R.P.G. Geraeds, D. Groenendijk, R. Ketelaar, R. Manger & T. Termaat 2008 Een actualisatie van de verspreiding van de Nederlandse libellen. Brachytron 11: 103-198

Collen, B., Ram, M., Dewhurst, N., Clausnitzer, V.J, Kalkman, V., Cumberlidge, N. and Baillie, J.E.M. 2008 Broadening the coverage of biodiversity assessments. In: J.-C. Vié, C. Hilton-Taylor and S.N. Stuart (eds). The 2008 Review of The IUCN Red List of Threatened Species. IUCN Gland, Switzerland. PDF

Darwall, W., K. Smith, D. Allen, M. Seddon, G. Mc Gregor Reid, V. Clausnitzer and V.J. Kalkman 2008 Freshwater biodiversity – a hidden resource under threat. In: J.-C. Vié, C. Hilton-Taylor and S.N. Stuart (eds.) The 2008 Review of The IUCN Red List of Threatened Species. IUCN, Gland, Switzerland. 

Dingemanse, N.J. & V.J. Kalkman 2008 Changing temperature regimes have advanced the phenology of Odonata in the Netherlands. Ecological Entomology 33: 1-9. 

Dyatlova, E.S. & V.J. Kalkman 2008 Massive migration of Aeshna mixta and Sympetrum meridionale in the Ukrainian Danube delta (Odonata-Anisoptera: Aeshnidae, Libellulidae). Entomologische Berichten 68: 188-190. PDF

Dyatlova, E.S. & V.J. Kalkman 2008 The Odonata of southwestern Ukraine, with emphasis on the species of the EU Habitats Directive. Libellula 27: 275-290.

Kalkman, V.J. 2008 Argiolestes in the Bismarck Archipelago and the Solomon Archipelago. Notes on Old World Megapodagrionidae 2 (Odonata). International Journal of Odonatology 11: 43-57.

Kalkman, V.J. 2008 Records of dragonflies from Borme, Star Mountains, Papua, Indonesia (Odonata). Entomologische Berichten 68: 45-52

Kalkman, V.J. 2008 Taxonomy, behaviour,and habitat of Mesopodagrion and Sinocnemis. Notes on Old World Megapodagrionidae 3. (Odonata). International Journal of Odonatology 11: 185-193. 

Kalkman, V.J. 2008 Two new dragonflies from Yapen and Biak, Papua (Irian Jaya), Indonesia (Odonata). Zoologische Mededelingen 82: 81-89. 

Kalkman, V.J., V. Clausnitzer, K.-D.B. Dijkstra, A.G. Orr, D.R. Paulson & J. van Tol 2008 Global diversity of dragonflies (Odonata) in freshwater. In: Balian, E., K. Martens, C. Lévêque & H. Segers (Editors). A global assessment of animal diversity in freshwater. Hydrobiologica 595: 351-363

Weide, M. van der & V.J. Kalkman 2008 Some new records of dragonflies from Oman. Agrion 12: 52-54. 


Kalkman, V.J. 2007 Argiolestes celebensis spec. nov. from Sulawesi, Indonesia (Odonata: Megapodagrionidae). Notes on old world Megapodagrionidae 1. Odonatologica 36: 295-299

Kalkman, V.J. 2007 Archboldargia scissorhandsi spec. nov from Papua, Indonesia (Odonata; Coenagrionidae). Odonatologica 36: 203-206.

Schouten, M.A., A. Barendregt, P.A. Verweij, V.J. Kalkman, R.J.M. Kleukers, H.J.R. Lenders & H.N. Siebel 2007 Representing biodiversity: spatial analysis of multi-taxon species composition in the Netherlands. In: Schouten, M.A. 2007. Patterns in biodiversity, spatial organisation of biodiversity in the Netherlands. Knag/Copernicus Institute, Utrecht.



Kalkman, V.J. 2006 Key to the dragonflies of Turkey, including species known from Greece, Bulgaria, Lebanon, Syria, the Trans-Caucasus and Iran. Brachytron 10: 3-82

Kalkman, V.J. 2006 Genus and species text of Epallage, Pyrrhosoma, Ceriagrion, Anax, Onychogomphus, Paragomphus, Lindenia, Orthetrum, Brachythemis, Selysiothemis. In: Dijkstra & Lewington, Fieldguide to the dragonflies of Britain and Europe.

Kalkman, V.J. & P. Hope 2006 Türkiye’nin Kizböcekleri. In: Eken, G., Bozdoğan, M. İsfendiyaroğlu, S., Kılıç, D.T., Lise, Y. (editörler) 2006. Türkiye’nin Önemli Doğa Alanları I. [Key Biodiversity Areas of Turkey] Doğa Derneği, Ankara. 473pp.

Kalkman, V.J. & R.M.J.C. Kleukers 2006 Collecties: onmisbaar voor  biodiversiteitsonderzoek. De Levende Natuur 107: 252-255. 

Kalkman, V.J. & W. Lopau 2006 Identification of Pyrrhosoma elisabethae with notes on its distribution and habitat (Odonata: Coenagrionidae). International Journal of Odonatology 9: 175-184. PDF

Kalkman, V.J. & B. Koese 2006 Herontdekking van een populatie van de Gewone bronlibel (Cordulegaster boltonii)  bij Venlo. Brachytron 9: 58-60

Kalkman, V.J. & G.J. van Pelt 2006 The distribution and flight period of the dragonflies of Turkey. Brachytron 10: 83-153

Kalkman, V.J. & G.J. van Pelt 2006 New records of rare or uncommon dragonflies in Turkey (Odonata). Brachytron 10: 154-162

Bouwman, J.H. & V.J. Kalkman 2006 Verspreiding van de libellen van de Habitatrichtlijn in Nederland. Brachytron 9: 3-13.



Kalkman, V.J. 2005 On the distribution of the genus Ceriagrion Selys in the Balkan, with C. georgfreyi Schmidt new for Europe. Libellula Supplement 6: 25-32. 

Reemer, R., F. Kok, R.H. de Bruyne, V.J. Kalkman & H. Turin 2005 Suitability of different groups of terrestrial invertebrates for assessment of heterogeneity of terrestrial parts of lowland floodplains. Archiv für Hydrobiologie 155: 289-303. 


Dijkstra, K.D.B. & V.J. Kalkman 2004 Een odonatologische exursie naar Zuid-Nederland, een halve eeuw later [An odonatological excursion to the southern Netherlands half a century later]. Entomologische Berichten 64: 157-161. 

Goudsmits, K. & V.J. Kalkman 2004 Zeldzame libellen in Nederland in 1999, 2000 en 2001, CWNO-mededeling 3 [Rare dragonflies in the Netherlands in 1999, 2000 and 2001, CWNO-reports 3]. Brachytron 8: 15-23.

Kalkman, V.J. 2004 Lindenia inkiti (Bartenev, 1929) a synonym of Lindenia tetraphylla(Vander Linden, 1825) (Anisoptera: Gomphidae). Notulae Odonatologicae 6(3): 34

Kalkman, V.J., A. Kop, M. Wasscher & G.J. van Pelt 2004 The dragonflies of the surroundings of lake Köyceğiz and the River Eşen, Muğla province, SW Turkey. Libellula supplement 5: 39-63. 

Kalkman, V.J., G.J. van Pelt, H.J. Dumont, A. Haritonov & M. Tailly 2004 Critical species of Odonata in Turkey, Iran and the Caucasus. In Clausnitzer, V. & R. Jödicke (eds), Guardians of the watershed. Global status of dragonflies: critical species, threat and conservation. International Journal of Odonatology 7: 139-147. 

Kalkman, V.J., W. Lopau & G.J. van Pelt 2004 Hitherto unpublished records of dragonflies from Turkey. – Libellula supplement 5: 65-166. 

Van Pelt, G.J. & V.J. Kalkman 2004 Research on dragonflies in Turkey: present status and future aims. – Libellula supplement 5: 167-192.


Kalkman, V.J. & N. Th. Wasscher 2003 Zeldzame libellen in 1998, CWNO-mededeling 2. Brachytron 7: 15-22

Kalkman, V.J., M. Wasscher & G.J. van Pelt 2003 An annotated checklist of the Odonata of Turkey. Odonatologica 32(3): 215-236


Kalkman, V.J. & J.T. Smit 2002 Platycnemis subdilitata Sel. new to the Canary Islands? (Zygoptera: Platycnemididae). Notulae-Odonatologicae 5 (10): 128.

Kalkman, V.J., G.A. van Duinen, H. Esselink & J.T. Kuper 2002 New records of Odonata from Estonia, with notes on breeding in the Baltic Sea and on species assemblages of raised bog systems. Notulae-Odonatologicae 5 (10): 120-125.

Nederlandse Vereniging voor Libellenstudie (Editors K.-D.B. Dijkstra, V.J. Kalkman, R. Ketelaar & M.J.T. van der Weide) 2002 De Nederlandse libellen (Odonata) [The Dutch dragonflies (Odonata)]. Nationaal Natuurhistorisch Museum Naturalis, KNNV Uitgeverij & European Invertebrate Survey, Leiden. 1-440.



Dijkstra, K.-D.B. & V.J. Kalkman 2001 Early spring records of Odonata from southern Turkey, with special reference to the sympatric occurrence of Crocothemis erythraea (Brullé) and C. servilia (Drury) (Anisoptera: Odonata). Notulae Odonatologicae, 5: 85-88. 

Kalkman, V.J. & N.J. Dingemanse 2001 Het voorkomen van de kleinvlekkige en grootvlekkige vorm van de Noordse glazenmaker (Aeshna subarctica) in Nederland [The occurence of the form interlineata and the typical form of Aeshna subarctica elisabethae in The Netherlands. Brachytron 5: 15-18


Kalkman, V.J. & K.-D.B. Dijkstra 2000 The dragonflies of the Bialowieza area, Poland and Belarus (Odonata). Opuscula zoologica fluminensia, 185: 1-19

Kalkman, V.J. 2000 Records on the dragonfly fauna of northwestern Albania (Odonata). Libellula, 19 (1-2): 107-111. 

Kalkman, V.J., P. Edelaar & M.A.A.M. van Trigt 2000 De voortplantingbiotoop van de Gevlekte glanslibel (Somatochlora flavomaculata) in Nederland [The breeding habitat of Somatochlora flavomaculata in the Netherlands. Brachytron 4: 28.


Kalkman, V.J. & R. Ketelaar 1999 [Interesting new records of Odonata in The Netherlands in 1998]. Nederlandse Faunistische Mededelingen, 8: 85-88.

Kalkman, V.J. 1999 [Migration of the hover fly Episyrphus balteatus on Falsterbo.]. Amoeba 73: 7-9.

Kalkman, V.J. 1999 De Danske guldesmede (Book review). Brachytron 3: 30.

Rutten, A. & V.J. Kalkman 1999 Eerste bewezen voortplanting van de Kempense heidelibel (Sympetrum depressiusculum (Selys)) in Nederland. [First evidence for succesfull reproduction of Sympetrum depressiusculum in the Netherlands]. Brachytron 3: 29-30.


Dijkstra, K.-D.B., D. Groenendijk & V.J. Kalkman 1998 Voorpost van het zuiden: de libellen van de Nedrelandse Sint Pietersberg [Odonata of the Dutch St. Pietersberg.]. Brachytron, 2: 23-27.

Zekhuis, M.J. & D. Tempelman [Eds] With contributions from: T. Bino, P. Defos du Rau, V.J. Kalkman, H. van Kleef, F. Willems, & E. van Winden 1998. Breeding birds of the Albanian wetlands, spring 1996 WIWO-Report 64: 1-143.


Dingemanse, N.J. & V.J. Kalkman 1997 Het onderscheid tussen Aeshna subarctica enAeshna juncea (L.) in Nederland [Separating adult Aeshna subarctica Walker from Aeshna juncea (L.) in the Netherlands]. Brachytron, 1 (2): 35-39


Ketelaar, R. & V.J. Kalkman 1995 [Observations on butterflies in Luxembourg]. Amoeba, 69 (3): 116-122.

Wasscher, M., R. Ketelaar, M. van der Weide, A. Stroo, V.J. Kalkman, N. Dingemanse, H. Inberg, I. Tieleman 1995 [Distribution status of Dutch dragonflies.]. Stichting European Invertebrate Survey-Nederland. Leiden. 1995: 1-80.


Kalkman, V.J. & A. Stroo 1993 [Manual Dutch Dragonfly mapping sheme.]. Jeugdbondslibellenproject, Utrecht.