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Steve Donovan

Prof. Dr. S.K. (Steve) Donovan, Senior researcher - Geology, Palaeozoic and Mesozoic invertebrates


Email: steve.donovan@naturalis.nl
Phone: +31 (0)71-56 87 642
Room number: C01.10, Darwinweg 2
“It all happened one Tuesday afternoon.”



D.Sc. 1994 Department of Earth Sciences, University of Liverpool, England.
Ph.D. 1983 Department of Geology, University of Liverpool, England.
B.Sc. 1980 Department of Geology, University of Manchester, England. Honours, Upper Second.



1983 Evolution and Biostratigraphy of Pelmatozoan Columnals from the Cambrian and Ordovician of Britain. Unpublished Ph.D. thesis, University of Liverpool, xx+557 pp.


Professional experience

2001-present Researcher of Mesozoic and Palaeozoic Macro-Invertebrates, and editor of Scripta Geologica Naturalis Biodiversity Center, Leiden
1998-2001 Keeper (=Head) of Palaeontology The Natural History Museum, London
1996‑1998 Professor of Palaeozoology, Department of Geography & Geology University of the West Indies, Jamaica
1994, 1995 Senior Research Fellow National Museum of Natural History, Smithsonian Institution, Washington, D.C., U.S.A
1992‑1996 Reader in Palaeozoology Department of Geology, University of the West Indies, Jamaica
1991 Reappointed by the University of the West Indies on indefinite tenure  
1989‑1992 Senior Lecturer Department of Geology, University of the West Indies, Jamaica
1986‑1989 Lecturer Department of Geology, University of the West Indies, Jamaica
1985‑1986 Higher Scientific Officer Natural Environment Research Council, Swindon, U.K.
1984 Palynology Technical Assistant Department of Geology, Trinity College, Dublin
1983‑1984 Royal Society Research Fellow Department of Geology, Trinity College, Dublin
1983 Research Assistant Department of Geology, University of Liverpool


Fellowships, awards and presidency

2013 Fellow, Geological Society of America.
2008 Linnean Medal for Zoology. Awarded by the Linnean Society of London.
1996 University of the West Indies Vice‑Chancellor's Award for Excellence in Research. US$5,000.
1994/5 Smithsonian Institution Senior Research Fellow at the National Museum of Natural History for 6 months (July 1994, April‑August 1995). US$14,104 (including allowances).
1992   Leverhulme Fellowship. UK,3,500 to undertake joint research in the UK with Professor C.R.C. Paul, University of Liverpool, on the biodiversity of early Palaeozoic and late Cenozoic carbonate environments.
1992 First Shell Distinguished Research Fellowship in Science. J$200,000 over 3 years to research the evolution and extinction of Caribbean Eocene‑Oligocene echinoids.
1983/4 Royal Society European Science Exchange Fellowship. Crinoid biostratigraphy and evolution in the Ordovician of Europe. Circa IR,13,000.
1979/80 President, University of Manchester Geological Society.



Research interest

My research background is broad. I have worked on the geology, palaeontology and stratigraphy of diverse groups and successions in northern Europe, the Caribbean, North America and elsewhere since 1980. 



Current research topics

Current research concerns the mid-Palaeozoic crinoids of northern Europe; Caribbean geology and palaeontology, particularly Jamaica and the Upper Oligocene of Antigua (the latter supported by the National Geographic Society); trace fossils; history of geology; and urban geology.


Apart from my colleagues at Naturalis, my principal collaborators include Drs Fiona Fearnhead and Tim Littlewood, and Mr David Lewis (all the Natural History Museum, London), Professor David Harper (Durham University), Dr John Jagt (Natuurhistorisch Museum Maastricht), Mr Roger W. Portell (Florida Museum of Natural History, Gainesville), Dr Donovan J. Blissett (University of the West Indies, Jamaica), Professors Ron K. Pickerill and David G. Keighley (both University of New Brunswick, fredericton) and Professor Trevor Jackson (University of the West Indies, Trinidad).



not applicatble

Supervision MSc Students

Recent M.Sc. projects that I have supervised include:

Baalsbergen, E. 2011. Terrestrial Arthropods from the Late Pleistocene of Jamaica. University of Leiden: 51 pp.

Kooijman, N. 2011. Fossil Echinoids from the Upper Oligocene Antigua Formation of Antigua, Lesser Antilles. University of Leiden: 32 pp.

Schelfhorst, R. 2013. The Echinodermata of the Pliocene of Curaçao. University of Leiden: 36 pp.

Ende, C. van den. 2013. Urchins on the Edge: A mixed environmental signal from the Eocene of Jamaica. University of Leiden: iv+39 pp.

Bouman, R.W. 2014. Biodiversity of Mississippian echinoderms from Derbyshire, UK. University of Leiden: 43 pp.

Available student projects

Public outreach

I am a regular contributor to the journal for amateurs Deposits (UK) in which I have written on a diversity of topics.


Recent peer reviewed publications: 

in press

Donovan, S.K. 2015 (in press). An ill wind: a gale, beachcombing and Gastrochaenolites Leymerie in peat, North Sea coast, the Netherlands. Bulletin of the Mizunami Fossil Museum41.

Donovan, S.K. 2014 (in press). Palaeoecology and taphonomy of a fossil ‘sea floor’, Mississippian (Carboniferous Limestone) of northern England. Mercian Geologist: 4 pp.

Donovan, S.K. 2014 (in press). When is a fossil not a fossil? When it is a trace fossil. Lethaia.

Donovan, S.K. 2014 (in press). History of palaeontology (version 2.0). In Encyclopedia of Life Sciences. John Wiley & Sons, Chichester.

Donovan, S.K. & Fearnhead, F.E. 2014 (in press). A dearth of diplobathrids (Crinoidea) from the type Devonian System, south-west England. Proceedings of the Yorkshire Geological Society60: 7 pp.

Donovan, S.K. & Fearnhead, F.E. 2014 (in press). The British Devonian Crinoidea. Part 1, introduction and Camerata. Monograph of the Palaeontographical Society168 (643): 1-55+i-viii, 15 pls.

Donovan, S.K. & Fearnhead, F.E. 2015 (in press). Exceptional fidelity of preservation in a reworked fossil, Chalk Drift, South London, England. Geological Journal: 3 pp.

Donovan, S.K., Jagt, J.W.M. & Goffings, L. 2014 (in press). Bored and burrowed: an unusual echinoid steinkern from the Type Maastrichtian (Upper Cretaceous, Belgium). Ichnos.

Donovan, S.K. & Novak, V. 2014 (in press). Site selectivity of predatory borings in Late Pliocene balanid barnacles from south-east Spain. Lethaia: 3 pp.

Donovan, S.K. & Portell, R.W. (in press). Invertebrate borings from the Eocene of Seven Rivers, parish of St. James, western Jamaica. In: Domning, D.P. & Portell, R.W. (eds), The Eocene Fossil Site of Seven Rivers, Jamaica: Geology, Paleontology, and Evolutionary and Biogeographic Implications. Springer, Heidelberg.



Collins, J.S.H., Donovan, S.K. & Mellish, C. 2014. An illustrated guide to the fossil barnacles (Cirripedia) from the Crags (Pilo-Pleistocene) of East Anglia. Proceedings of the Geologists’ Association125: 215-226.

Donovan, S.K. 2014. Pioneers of Jamaican geology and the Jamaica Group of the Geologists’ Association (1955-1959). Proceedings of the Geologists’ Association125: 131-138.

Donovan, S.K. 2014. Discussion on “The Gramscatho Basin, south Cornwall, UK: Devonian active margin successions” [Proceedings of the Geologists’ Association 122 (2011) 568-615]. Proceedings of the Geologists’ Association125: 255.

Donovan, S.K. 2014. Enigmatic branching structures within Upper Devonian crinoids, north Devon, UK. Lethaia47: 151-152.

Donovan, S.K. 2014. The trace fossil Teredolites longissimus Kelly and Bromley, 1984, from the Eocene of Curaçao. Caribbean Journal of Science47 (for 2013): 181-185.

Donovan, S.K. 2014. How Not to Alienate your Editor: Some Everyday Aspects of Academic Publishing. Lambert Academic Publishing, Saarbrücken: 107 pp.

Donovan, S.K., Ausich, W.I., Wilson, M.A. & Peter, M.E. 2014. A Middle Ordovician crinoid from the beach gravels of Ristna Cape, Hiiumaa Island, Estonia. Proceedings of the Geologists’ Association125: 96-98.

Donovan, S.K., Cotton, L., Ende, Conrad van den, Scognamiglio, G. & Zittersteijn, M. 2014. Taphonomic significance of a dense infestation of Ensis americanus (Binney) by Balanus crenatus Brugière, North Sea. Palaios28 (for 2013): 837-838.

Donovan, S.K., Harper, D.A.T., Jackson, T.A. & Portell, R.W. 2014. A note on a coastal natural bridge in Antigua, West Indies. Cave & Karst Science40 (for 2013): 105-108.

Donovan, S.K., Harper, D.A.T., Portell, R.W. & Renema, W. 2014. Neoichnology and implications for stratigraphy of reworked Upper Oligocene oysters, Antigua, West Indies. Proceedings of the Geologists’ Association125: 99-106.

Donovan, S.K. & Isted, J. 2014. Teredolites Leymerie in the Lower Greensand Group (Cretaceous) of the Isle of Wight and the problematic ichnology of reworked clasts. Proceedings of the Geologists’ Association125: 252-254.

Donovan, S.K. & Jagt, J.W.M. 2014. Ichnology of Late Cretaceous echinoids from the Maastrichtian type area (The Netherlands, Belgium) – 3. Podichnus Bromley and Surlyk and a crinoid attachment on the echinoid Echinocorys Leske from the Lixhe area, Belgium. Bulletin of the Mizunami Fossil Museum40: 75-78.

Donovan, S.K. & Lewis, D.N. 2014. An illustrated guide to the Silurian (Ludlow) crinoids of the southern Lake District. Proceedings of the Cumberland Geological Society8 (for 2012-2013): 180-191.

Donovan, S.K. & Portell, R.W. 2014. Fossil echinoids from the Upper Pliocene Hopegate Formation of north central Jamaica. Caribbean Journal of Science47 (for 2013): 125-139.

Ende, C. Van den & Donovan, S.K. 2014. Internal disorder: post-mortem burrows within the tests of the holasteroid echinoid Echinocorys scutata (Leske) (Upper Cretaceous to Paleocene). Ichnos21: 73-75.

Jagt, J.W.M., Thuy, B., Donovan, S.K., Stöhr, S., Portell, R.W., Pickerill, R.K., Harper, D.A.T., Lindsay, W. & Jackson, T.A. 2014. A starfish bed in the Middle Miocene Grand Bay Formation of Carriacou, The Grenadines (West Indies). Geological Magazine151: 381-393.

Ouwendijk, M., Hoek Ostende, L.W. van den & Donovan, S.K. 2014. Fossil bats from the Late Pleistocene Red Hills Road Cave, Jamaica. Caribbean Journal of Science47 (for 2013): 284-290.



Baalbergen, E. & Donovan, S.K. 2013. Terrestrial arthropods from the Late Pleistocene of Jamaica: systematics, palaeoecology and taphonomy. Geological Journal, 48: 628-645.

Donovan, S.K. 2013. A Recent example of the boring Gastrochaenolites lapidicus Kelly and Bromley and its producing organism in north Norfolk, eastern England. Bulletin of the Mizunami Fossil Museum39: 69-71.

Donovan, S.K. 2013. Misinterpreting by localism: transposing European geology and tectonics onto Jamaica and the Antilles. Proceedings of the Geologists’ Association124: 530-535.

Donovan, S.K. 2013. Curiouser and curiouser: more on reworked Echinocorys (Echinoidea; Late Cretaceous) on the beaches of north Norfolk, eastern England. Swiss Journal of Palaeontology132: 1-4.

Donovan, S.K. 2013. Giant crinoid stems from the Lower Carboniferous (Mississippian) of northwest England. Proceedings of the Yorkshire Geological Society59: 211-218.

Donovan, S.K. 2013. A distinctive bioglyph and its producer: Recent Gastrochaenolites Leymerie in a peat pebble, North Sea coast of the Netherlands. Ichnos20: 109-111.

Donovan, S.K. 2013. The cladid crinoid Cupressocrinites Goldfuss in the Devonian of SW England. Proceedings of the Yorkshire Geological Society59: 255-259.

Donovan, S.K., Baalbergen, E., Ouwendijk, M., Paul, C.R.C. & Hoek Ostende, L.W. van den. 2013. Review and prospectus of the Late Pleistocene fauna of the Red Hills Road Cave, Jamaica. Cave and Karst Science40: 79-86.

Donovan, S.K. & Hoek Ostende, L.W. van den. 2013. Roamerella, a new name for the genus Roemerella Akopyan in Akopyan et al., 1990 (Mollusca, Gastropoda; Upper Cretaceous), preoccupied by Roemerella Hall and Clarke, 1890 (Brachiopoda, Lingulata; Devonian). Cretaceous Research40: 1-2.

Donovan, S.K. & Jagt, J.W.M. 2013. Site selectivity of the pit Oichnus excavatus Donovan and Jagt infesting Hemipneustes striatoradiatus (Leske) (Echinoidea) in the Type Maastrichtian (Upper Cretaceous, The Netherlands). Ichnos20: 112-115.

Donovan, S.K. & Jagt, J.W.M. 2013. Aspects of clavate borings in the type Maastrichtian (Upper Cretaceous) of the Netherlands and Belgium. Netherlands Journal of Geosciences92: 133-143.

Donovan, S.K. & Jagt, J.W.M. 2013. Rogerella isp. infesting the pore pairs of Hemipneustes striatoradiatus (Leske) (Echinoidea: Upper Cretaceous, Belgium). Ichnos20: 153-156.

Donovan, S.K. & Paul, C.R.C. 2013. Late Pleistocene land snails from ‘red bed’ deposits, Round Hill, south central Jamaica. Alcheringa37: 273-284.

Donovan, S.K. & Pawson, D.J. 2013. An intersection in time and space: significance of modern invertebrate borings in Upper Cretaceous echinoids. Ichnos20: 81-87.

Donovan, S.K. & Pickerill, R.K. 2013. On marls and marlstones. Bulletin of the Mizunami Fossil Museum39: 127-128.

Donovan, S.K. & Riley, M. 2013. The importance of labels to specimens: an example from the Sedgwick Museum. The Geological Curator9: 509-514.

Donovan, S.K., Waveren, I.M. van. & Portell, R.W. 2013. Island slopes and jumbled shell beds. Journal of the Geological Society170: 527-534.

Donovan, S.K. & Webster, G.D. 2013. Platyceratid gastropod infestations of Neoplatycrinus Wanner (Crinoidea) from the Permian of West Timor: speculations on thecal modifications. Proceedings of the Geologists’ Association124: 988-993.

Fearnhead, F.E., Donovan, S.K. & Ewin, T.A.M. 2013. Palaeobiology of Floricolumnus (col.) girvanensis Donovan & Clark (Crinoidea: Silurian) from the Girvan district, Ayrshire. Scottish Journal of Geology49: 1-7.



Collins, J.S.H. & Donovan, S.K. 2012. Pleistocene decapod crustaceans of eastern Jamaica. Caribbean Journal of Science46: 133-142.

Donovan, S.K. 2012. Stanley Westhead and the Lower Carboniferous (Mississippian) crinoids of the Clitheroe area, Lancashire. Proceedings of the Yorkshire Geological Society59: 15-20.

Donovan, S.K. 2012. Comment: Taxonomic diversity structure of Silurian crinoids: stability versus dynamism. Annales de Paléontologie98: 153-155.

Donovan, S.K. 2012. Was autotomy a pervasive adaptation of the crinoid stalk during the Palaeozoic? Geology40: 867-870.

Donovan, S.K. 2012. Taphonomy and significance of rare Chalk (Late Cretaceous) echinoderms preserved as beach clasts, north Norfolk, UK. Proceedings of the Yorkshire Geological Society59: 109-113.

Donovan, S.K. 2012. Notes on Lower Devonian crinoids in the collections of the British Geological Survey, Keyworth. Proceedings of the Yorkshire Geological Society59: 115-120.

Donovan, S.K. 2012. An unusual accumulation of crinoids from the Silurian of Howgill Fells, Cumbria, UK. Proceedings of the Yorkshire Geological Society59: 121-123.

Donovan, S.K., Harper, D.A.T., Zhan Renbin, Stemmerik, L., Liu Jianbo & Stouge, S. 2012. A primitive cladid crinoid from the Jiacun Group, Tibet (Darriwilian, Middle Ordovician). Geological Journal47: 653-660.

Donovan, S.K. & Hoek Ostende, L.W. van den. 2012. Neoextrania, a replacement name for Extrania Sun & Wang, 1985 (Ostracoda), preoccupied by Extrania Qian in Qiu et al., 1983 (Trilobita). Alcheringa36: 423.

Donovan, S.K. & Jagt, J.W.M. 2012. Dentaliids (Mollusca, Scaphopoda) from the type Maastrichtian, the Netherlands and Belgium. In Jagt, J.W.M., Donovan, S.K. & Jagt-Yazykova, E.A. (eds), Fossils of the type Maastrichtian. Scripta Geologica Special Issue, 8: 83-91.

Donovan, S.K., Renema, W., Pinnington, C.A. & Veltkamp, C.J. 2012. Significance of diadematid echinoid ossicles in micropalaeontological samples, Miocene-Pliocene of Indonesia. Alcheringa36: 99-105.

Donovan, S.K. & Schelfhorst, R. 2012. A Pliocene asteroid from the Netherlands Antilles. Bulletin of the Mizunami Fossil Museum38: 75-77.

Donovan, S.K., Widdison, R.E., Lewis, D.N. & Fearnhead, F.E. 2012. The British Silurian Crinoidea. Part 3, addendum to parts 1 and 2, Camerata and columnals. Monograph of the Palaeontographical Society, London, 166 (638): 135-259, pls 37-62.

Jagt, J.W.M., Donovan, S.K. & Deckers, M.J.M. 2012. Clustered bourgueticrinid crinoid holdfasts on late Maastrichtian echinoids from northeast Belgium and southeast Netherlands. In: Kroh, A. & Reich, M. (eds), Echinoderm Research 2010: Proceedings of the Seventh European Echinoderm Conference, Göttingen, Germany, 2-9 October 2010. Zoosymposia, 7: 81-90.

Jagt, J.W.M., Donovan, S.K. & Jagt-Yazykova, E.A. (eds) 2012. Fossils of the type Maastrichtian (Part 1). Scripta Geologica Special Issue, 8: 142 pp.

Jagt, J.W.M., Donovan, S.K. & Jagt-Yazykova, E.A. 2012. Fossils of the type Maastrichtian. In Jagt, J.W.M., Donovan, S.K. & Jagt-Yazykova, E.A. (eds), Fossils of the type Maastrichtian. Scripta Geologica Special Issue, 8: 1-4.

Taylor, P.D. & Donovan, S.K. 2012. A bryozoan from the Upper Pliocene Hopegate Formation of north central Jamaica. Caribbean Journal of Earth Science44: 5-8.

Webster, G.D. & Donovan, S.K. 2012. Revision of two species of ?Ulocrinus and a new pelecocrinid from West Timor. Palaeoworld21: 108-115.

Webster, G.D. & Donovan, S.K. 2012. Before the extinction – Permian platyceratid gastropods attached to platycrinitid crinoids and an abnormal four-rayed Platycrinites s.s. wachsmuthi (Wanner) from West Timor. Palaeoworld21: 153-159.



Andrew, C., Howe, P., Paul, C.R.C. & Donovan, S.K. 2011. Epifaunal worm tubes on Lower Jurassic (Lower Lias) ammonites from Dorset. Proceedings of the Geologists’ Association122: 34-46.

Donovan, S.K. 2011. 150 years of managing national geological assets: celebrating the sesquicentennial of the first Official Geological Survey in Jamaica: comment. Caribbean Journal of Earth Science42: 17-19.

Donovan, S.K. 2011. Post-mortem encrustation of the alien bivalve Ensis americanus (Binney) by the barnacle Balanus crenatus Brugière in the North Sea. Palaios26: 665-668.

Donovan, S.K. 2011. A field guide to the Cretaceous and Cenozoic fossil decapod crustacean localities of Jamaica. Caribbean Journal of Science46 (for 2010): 39-53.

Donovan, S.K. 2011. Writing about geology. Geological Journal46: 374-379.

Donovan, S.K. 2011. Spineless displays or why inaccurate restorations of fossil invertebrates discredit our museums. Geological Curator9: 279-284.

Donovan, S.K. (ed.). 2011. Islands: Palaeontology, Geology and Tectonics. Proceedings of the Lyell Meeting 2011, Burlington House, London, Monday, 24th October. Scripta Geologica, 142: 1-27.

Donovan, S.K. 2011. The poorly illustrated crinoid. Lethaia44: 125-135.

Donovan, S.K. 2011. Aspects of ichnology of Chalk and sandstone clasts from the beach at Overstrand, north Norfolk. Bulletin of the Geological Society of Norfolk60 (for 2010): 37-45.

Donovan, S.K. 2011. Crinoids from the type area of the Costonian (Upper Ordovician), south Shropshire, England. Bulletin of the Mizunami Fossil Museum37: 181-184.

Donovan, S.K. 2011. The Recent boring Gastrochaenolites ornatus Kelly & Bromley, 1984, in a Chalk cobble from Cromer, England. Bulletin of the Mizunami Fossil Museum37: 185-188.

Donovan, S.K. & Birtle, M. 2011. The cladid crinoid Rhabdocrinus Wright from the Namurian of Co. Durham, UK. Proceedings of the Yorkshire Geological Society58: 167-171.

Donovan, S.K., Fearnhead, F.E., Clarkson, E.N.K. & Donovan, M. 2011. Aspects of echinoderm palaeontology of the North Esk Inlier, Scotland (Silurian, Llandovery, Telychian). Palaeontology54: 241-252.

Donovan, S.K. & Harper, D.A.T. (eds). 2011. Early Palaeozoic Ecosystems, Environments and EvolutionGeological Journal46: 395-512.

Donovan, S.K. & Harper, D.A.T. 2011. Early Palaeozoic ecosystems, environments and evolution: a synopsis. Geological Journal46: 395-396.

Donovan, S.K., Harper, D.A.T. & Marshall, J.D. 2011. Patrick John Brenchley (1936-2011). Geological Journal46: 397-404.

Donovan, S.K. & Hoek Ostende, L.W. van den. 2011. Rijckholtia, a new name for the genus Ryckholtia Šnajdr, 1980 (Arthropoda, Trilobita), preoccupied by Ryckholtia Fritsch, 1910 (Mollusca, Gastropoda). Journal of Paleontology85: 804.

Donovan, S.K., Jagt, J.W.M. & Lewis, D.N. 2011. Notes on some trace fossils and other parataxa from the Maastrichtian type area, southeast Netherlands and northeast Belgium. Netherlands Journal of Geosciences90: 99-109.

Donovan, S.K. & Lewis, D.N. 2011. Fossil echinoderms from the Mississippian (Lower Carboniferous) of the Clitheroe district. In Kabrna, P.N. (ed.), Carboniferous Geology: Bowland Fells to Pendle Hill: 55-96. Craven & Pendle Geological Society, UK.

Donovan, S.K. & Lewis, D.N. 2011. Strange taphonomy: Late Cretaceous Echinocorys (Echinoidea) as a hard substrate in a modern shallow marine environment. Swiss Journal of Palaeontology130: 43-51.

Donovan, S.K. & Mellish, C.J.T. 2011. Presentation of the 2010 Harrell L. Strimple Award of the Paleontological Society to J.S.H. “Joe” Collins. Journal of Paleontology85: 1017-1018.

Donovan, S.K., Miller, C.G., Sansom, I.J., Howard, A.P. & Schreurs, J. 2011. A Laurentian Iocrinus Hall (Crinoidea, Disparida) in the Dapingian or Darriwilian (Middle Ordovician, Arenig) of Oman. Palaeontology54: 525-533.

Donovan, S.K. & Paul, C.R.C. 2011. Land snails from the late Pleistocene lithified sand dunes of Great Pedro Bluff, southwest Jamaica. Caribbean Journal of Science46 (for 2010): 1-11.

Donovan, S.K. & Riley, M. 2011. What's in a (wrong) name? Thoughts on the true utility of electronic museum catalogues. The Geological Curator9: 357-362.

Donovan, S.K., Sutton, M.D. & Sigwart, J.D. 2011. The last meal of the Late Ordovician mollusc ‘Helminthochiton’ thraivensis Reed, 1911, from the Lady Burn Starfish Beds, southwest Scotland. Geological Journal46: 451-463.

Harper, D.A.T., Zhan Renbin, Stemmerik, L., Liu Jianbo, Donovan, S.K. & Stouge, S. 2011. Ordovician on the roof of the World: macro- and microfaunas from tropical carbonates in Tibet. In: Gutiérrez-Marco, J.C., Rábano, I. & García-Bellido, D. (eds), Ordovician of the World. Cuadernos del Museo Geominero, 14: 215-220.

Paul, C.R.C. & Donovan, S.K. 2011. A review of the British Silurian cystoids. Geological Journal46: 434-450.



Andrew, C., Howe, P., Paul, C.R.C. & Donovan, S.K. 2010. Fatally bitten ammonites from the lower Lias Group (Lower Jurassic) of Lyme Regis, Dorset. Proceedings of the Yorkshire Geological Society58: 81-94.

Collins, J.S.H., Donovan, S.K., Stemann, T.A. & Blissett, D.J. 2010. Crustaceans of the Upper Miocene August Town Formation of eastern Jamaica. Scripta Geologica Special Issue7: 177-183.

Donovan, S.K. 2010. A Derbyshire screwstone (Mississippian) from the beach at Overstrand, Norfolk, eastern England. Scripta Geologica Special Issue7: 43-52.

Donovan, S.K. 2010. Blastoids, carpoids, crinoids, cystoids, edrioblastoids, eocrinoids, helicoplacoids. In MacLeod, N. (ed.), PaleoBase: Macrofossils 3.0. Wiley-Blackwell and The Natural History Museum, Oxford and London. [CD-ROM.]

Donovan, S.K. 2010. Cruziana and Rusophycus: trace fossils produced by trilobites ... in some cases? Lethaia43: 283-284.

Donovan, S.K. (ed.), with contributions by Chubb, L.J. and Williams, J.B. 2010. Jamaican Rock Stars 1823-1971: The Geologists who Explored Jamaica. Geological Society of America Memoir, 205: 108 pp.

Donovan, S.K. 2010. Jamaican rock stars. Geological Society of America Memoir205: 1-8.

Donovan, S.K. 2010. R.T. Hill (1858-1941) and “The geology and physical geography of Jamaica: study of a type of Antillean development” (1899). Geological Society of America Memoir205: 47-57.

Donovan, S.K. 2010. Three points of view: Wendell P. Woodring (1891-1983), Charles A. Matley (1866-1947), Charles T. Trechmann (1884-1964), and Jamaican geology in the 1920s and 1930s. Geological Society of America Memoir205: 59-78.

Donovan, S.K. 2010. An appreciation of Lawrence John Chubb (1887-1971), stratigrapher, educator, and historian of Jamaican geology. Geological Society of America Memoir205: 103-108.

Donovan, S.K. 2010. Where is Glasgow? Clarifying a Jamaican locality after 85 years. Bulletin of the Mizunami Fossil Museum36: 127-128.

Donovan, S.K., Blissett, D.J. & Jackson, T.A. 2010. Reworked fossils, ichnology and palaeoecology: an example from the Neogene of Jamaica. Lethaia43: 441-444.

Donovan, S.K. & Ewin, T.A.M. 2010. Crinoid roots from the Upper Devonian of north Devon: morphology, function and systematics. Proceedings of the Yorkshire Geological Society58: 15-20.

Donovan, S.K. & Harper, D.A.T. 2010. Nurse logs and nurse crinoids? A palaeobotanical concept applied to fossil crinoids. Lethaia43: 591-592.

Donovan, S.K. & Hoek Ostende, L.W. van den (eds). 2010. Palaeontological papers in honour of Dr. Cor F. Winkler Prins. Scripta Geologica Special Issue, 7: 183 pp.

Donovan, S.K., Jagt, J.W.M. & Dols, P.P.M.A. 2010. Ichnology of Late Cretaceous echinoids from the Maastrichtian type area (The Netherlands, Belgium) - 2. A pentagonal attachment scar on Echinocorys gr. conoidea (Goldfuss). Bulletin of the Mizunami Fossil Museum36: 51-53.

Donovan, S.K. & Lewis, D.N. 2010. Aspects of crinoid palaeontology, Much Wenlock Limestone Formation, Wenlock Edge, Shropshire (Silurian). Proceedings of the Yorkshire Geological Society58: 9-14.

Donovan, S.K. & Lewis, D.N. 2010. Notes on a Chalk pebble from Overstrand: ancient and modern sponge borings meet on a Norfolk beach. Bulletin of the Geological Society of Norfolk59 (for 2009): 3-9.

Donovan, S.K., Renema, W. & Lewis, D.N. 2010. A new species of Goniocidaris Desor (Echinoidea, Cidaroida) from the middle Miocene of Java. Alcheringa34: 87-95.

Donovan, S.K., Sutton, M.D. & Sigwart, J.D. 2010. Crinoids for lunch? An unexpected biotic interaction from the Upper Ordovician of Scotland. Geology38: 935-938.

Donovan, S.K., Widdison, R.E., Lewis, D.N. & Fearnhead, F.E. 2010. The British Silurian Crinoidea. Part 2, addendum to Part 1 and Cladida. Monograph of the Palaeontographical Society164: 47-133, pls 7-36.

Hoek Ostende, L.W. van den & Donovan, S.K. 2010. Cor Winkler Prins, the silent force. Scripta Geologica Special Issue7: 1-11.



Callomon, J.H., Donovan, S.K. & Hoek Ostende, L.W. van den. 2009. Zenostephanus, a new name for the genus Xenostephanus Arkell and Callomon, 1963 (Mollusca, Cephalopoda), preoccupied by Xenostephanus Simpson, Minoprio and Patterson, 1962 (Mammalia). Palaeontology52: 671-672.

Collins, J.S.H., Donovan, S.K. & Stemann, T.A. 2009. Fossil Crustacea of the Late Pleistocene Port Morant Formation, west Port Morant Harbour, southeastern Jamaica. Scripta Geologica138: 23-53.

Collins, J.S.H., Mitchell, S.F. & Donovan, S.K. 2009. A new species of land crab, Sesarma Say, 1817 (Decapoda, Brachyura), from the Pleistocene of Jamaica. Scripta Geologica138: 11-21.

Collins, J.S.H., Portell, R.W. & Donovan, S.K. 2009. Decapod crustaceans from the Neogene of the Caribbean: diversity, distribution and prospectus. Scripta Geologica138: 55-111.

Crow, M.J., Waveren, I.M. van & Donovan, S.K. 2009. Tobler’s oyster and the age of the Tabir Formation, Jambi Province, central Sumatra. Geological Journal., 44: 117-121.

Donovan, S.K. (ed.). 2009. Cenozoic crustaceans of the East and West Indies. Scripta Geologica, 138: 111 pp.

Donovan, S.K. 2009. John Phillips and the British Silurian Crinoidea: 170 years of monographic endeavour. Proceedings of the Yorkshire Geological Society57: 155-164.

Donovan, S.K. 2009. The camerate crinoid Eucalyptocrinites sp. from the Silurian(?) of Scotland. Scottish Journal of Geology45: 89-92.

Donovan, S.K., Fearnhead, F.E. & Clarkson, E.N.K. 2009. The ichnofossils Gordia Emmons and Chondrites von Sternberg from the Deerhope Formation, North Esk Inlier (Silurian, Llandovery). Scottish Journal of Geology45: 83-87.

Donovan, S.K., Harper, D.A.T. & Rasmussen, J.A. 2009. Taphonomy of logs bored with Teredolites longissimus Kelly and Bromley in the Danian (Lower Paleocene) of West Greenland. Ichnos16: 186-192.

Donovan, S.K. & Hoek Ostende, L.W. van den. 2009. What’s in a name? Or a namer’s name? A reply to Dubois (2008). Biological Journal of the Linnean Society96: 709-711.

Donovan, S.K. & Jakobsen, S.L. 2009. Comment on “The ‘living-fossil’ community of the cyrtocrinid Cyathidium foresti and the deep-sea oyster Neopycnodonte zibrowii (Azores Archipelago)” by M. Wisshak, C. Neumann, J. Jakobsen and A. Freiwald [Palaeogeography, Palaeoclimatology, Palaeoecology 271 (2009), 77-83]. Palaeogeography, Palaeoclimatology, Palaeoecology279: 233-234.

Donovan, S.K. & Lewis, D.N. 2009. The mid-Palaeozoic camerate crinoid Scyphocrinites Zenker in southwest England. Bulletin of the Mizunami Fossil Museum35: 97-100.

Donovan, S.K. & Lewis, D.N. 2009. Paleontological implications of multiple genital pores in the apical system of Eucidaris tribuloides (Lamarck), Recent of Jamaica. Caribbean Journal of Science45: 20-24.

Donovan, S.K., Lewis, D.N., Fearnhead, F.E. & Widdison, R.E. 2009. The British Silurian Crinoidea. Part 1, introduction and Disparida. Monographs of the Palaeontographical Society, London, 163 (632): 1-45, pls 1-6.

Donovan, S.K. & Portell, R.W. 2009. Reply to discussion of Campanile trevorjacksoni sp. nov. (Mollusca: Gastropoda) from the Eocene of Jamaica - at last, a name for the first fossil used in intercontinental biostratigraphic correlation (de la Beche 1827). Geological Journal44: 497-499.

Jagt, J.W.M., Neumann, C. & Donovan, S.K. 2009. Petroxestes altera, a new bioerosional trace fossil from the upper Maastrichtian of northeast Belgium. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique: Sciences de la Terre79: 137-145.

Meijer, H.J.M., Donovan, S.K. & Renema, W. 2009. Major Dutch collections of Permian fossils from Timor amalgamated. Journal of Paleontology83: p. 313.



Donovan, S.K. (ed.) 2008. Crustal and biotic evolution of the Caribbean plate. Geological Journal, 43 (5): 521-619.

Donovan, S.K. 2008. Professor Trevor A. Jackson. Geological Journal43: 521-524.

Donovan, S.K. 2008. The ‘Forbidden Theory of Mountain Uplift’ of Charles Taylor Trechmann (1884-1964): a tectonic theory of the 1950s in context. Geological Journal43: 605-619.

Donovan, S.K., Jagt, J.W.M. & Lewis, D.N. 2008. Ichnology of Late Cretaceous echinoids from the Maastrichtian type area (The Netherlands, Belgium) - 1. A healed puncture wound in Hemipneustes striatoradiatus (Leske). Bulletin of the Mizunami Fossil Museum34: 73-76.

Donovan, S.K. & Lewis, D.N. 2008. A crinoid crown from the Wenlock (Silurian) of Coalbrookdale, Shropshire, England. Scripta Geologica136: 63-72.

Donovan, S.K. & Lewis, D.N. 2008. A conundrum from the Llandovery (Lower Silurian) of Devil’s Dingle, Shropshire. Proceedings of the Yorkshire Geological Society57: 75-78.

Donovan, S.K., Lewis, D.N., Crabb, P. & Widdison, R.E. 2008. A field guide to the Silurian Echinodermata of the British Isles: Part 2 - Crinoidea, minor groups and discussion. Proceedings of the Yorkshire Geological Society57: 29-60.

Donovan, S.K., Lewis, D.N. & Veltkamp, C.J. 2008. The cladid crinoid Gissocrinus Angelin from Wenlock Edge, Shropshire (Silurian). Proceedings of the Yorkshire Geological Society57: 69-74.

Donovan, S.K., Lewis, D.N., Widdison, R.E. & Fearnhead, F.E. 2008. Ever since Ramsbottom: Silurian crinoids of the British Isles since 1954. In Ausich, W.I. & Webster, G.D. (eds), Echinoderm Paleobiology: 330-345. Indiana University Press, Bloomington.

Donovan, S.K. & Pawson, D.L. 2008. A new species of the sessile crinoid Holopus d’Orbigny from the tropical western Atlantic, with comments on holopodid ecology (Echinodermata: Crinoidea: Holopodidae). Zootaxa1717: 31-38.

Donovan, S.K. & Pickerill, R.K. 2008. The Paleogene Richmond Formation of Jamaica: not an impact-related succession. Scripta Geologica136: 107-111.

Jackson, T.A., Scott, P.W., Donovan, S.K., Pickerill, R.K., Portell, R.W. & Harper, D.A.T. 2008. The volcaniclastic turbidites of the Grand Bay Formation, Carriacou, Grenadines, Lesser Antilles. Caribbean Journal of Science44: 116-124.

Portell, R.W. & Donovan, S.K. 2008. Campanile trevorjacksoni sp. nov. (Mollusca: Gastropoda) from the Eocene of Jamaica: at last, a name for the first fossil used in intercontinental biostratigraphic correlation (de la Beche 1827). Geological Journal43: 542-551.

Portell, R.W., Hubbell, G., Donovan, S.K., Green, J.L., Harper, D.A.T. & Pickerill, R.K. 2008. Miocene sharks in the Kendeace and Grand Bay formations of Carriacou, the Grenadines, Lesser Antilles. Caribbean Journal of Science44: 279-286.

Stemann, T.A. & Donovan, S.K. 2008. Paleo-channel or sinkhole? Reinterpretation of an enigmatic structure near Discovery Bay, north central Jamaica. Caribbean Journal of Science44: 242-245.



Bruijn, H. de, Hoek Ostende, L.W. van den & Donovan, S.K. 2007. Mirrabella, a new name for the genus Mirabella De Bruijn et al. 1987 (Mammalia), preoccupied by Mirabella Emeljanov 1982 (Insecta). Contributions to Zoology76: 279-280.

Ciampaglio, C.N., Donovan, S.K. & Weaver, P.G. 2007. A new bourgueticrinid (Crinoidea) from the Castle Hayne Formation (Eocene) of southeastern North Carolina, USA. Swiss Journal of Geoscience100: 243-249.

Collins, J.S.H. & Donovan, S.K. 2007. New decapod crustaceans from the Paleogene of Jamaica. Bulletin of the Mizunami Fossil Museum33 (for 2006): 59-65.

Donovan, S.K. 2007. H.L. ‘Hal’ Dixon (1941-2005) and the fossil echinoids of Jamaica. Caribbean Journal of Science43: 279-282.

Donovan, S.K. 2007. A cautionary tale: razor shells, acorn barnacles and palaeoecology. Palaeontology50: 1479-1484.

Donovan, S.K. 2007. No Echinolampas marker band in the Miocene of Jamaica? Supporting evidence from Poland. Caribbean Journal of Science43: 147-150.

Donovan, S.K. 2007. Collectors, curators and the fossil record: a reply to Hope (2004). Scottish Journal of Geology43: 75-77.

Donovan, S.K., Fearnhead, F.E. & Lewis, D.N. 2007. Crinoids. In Clarkson, E.N.K., Harper, D.A.T., Taylor, C.M. & Anderson, L.I. (eds), Silurian Fossils of the Pentland Hills, Scotland. Palaeontological Association Field Guides to Fossils, 11: 172-180.

Donovan, S.K., Fearnhead, F.E. & Underwood, C.J. 2007. Recent borings in limestone cobbles from Marloes Bay, southwest Wales. Lethaia., 40: 233-236.

Donovan, S.K. & Harper, D.A.T. 2007. Rare borings in Pleistocene brachiopods from Jamaica and Barbados. Caribbean Journal of Science43: 59-64.

Donovan, S.K., Harper, D.A.T. & Håkansson, E. 2007. The root of the problem: palaeoecology of distinctive crinoid attachment structures from the Silurian (Wenlock) of Gotland. Lethaia40: 313-320.

Donovan, S.K., Portell, R.W. & Domning, D.P. 2007. Contrasting patterns and mechanisms of extinction during the Eocene-Oligocene transition in Jamaica. In Renema, W. (ed.), Biogeography, Time and Place: Distributions, Barriers and Islands: 247-273. Springer, Dordrecht.

Donovan, S.K. & Stemann, T.A. 2007. Where have all the sponges gone? The significance of spicules from the Upper Pliocene Hopegate Formation of Jamaica. Caribbean Journal of Science42 (for 2006): 239-243.

Fearnhead, F.E. & Donovan, S.K. 2007. A cladid crinoid (Echinodermata) from the Llandovery (Lower Silurian) of the Girvan district, SW Scotland. Scottish Journal of Geology43: 79-82. [2007. Corrigendum. Scottish Journal of Geology43: 180.]

Fearnhead, F.E. & Donovan, S.K. 2007. New crinoids (Echinodermata) from the Llandovery (Lower Silurian) of the British Isles. Palaeontology50: 905-915.

Fearnhead, F.E. & Donovan, S.K. 2007. A robust crinoid from the Llandovery (Lower Silurian) of Norbury, Shropshire: systematics, palaeoecology and taphonomy. Proceedings of the Geologists= Association118: 339-345.

Harper, D.A.T. & Donovan, S.K. 2007. Fossil brachiopods from the Pleistocene of the Antilles. Scripta Geologica135: 213-239.

Lewis, D.N. & Donovan, S.K. 2007. A standardized method of describing fossils, using Echinoidea as an example. Scripta Geologica134: 109-118.

Lewis, D.N., Donovan, S.K., Crabb, P. & Gladwell, D.J. 2007. A field guide to the Silurian Echinodermata of the British Isles: Part 1 - Eleutherozoa and Rhombifera. Scripta Geologica134: 27-59.

MacGillivray, C.M.I. (prepared for publication by Donovan, S.K.). 2007. A Relative Potential Erosion Detection (PED) model for the upper Buff Bay catchment, parish of Portland, Jamaica: A geographical information system application. Scripta Geologica Special Issue6: 202 pp.



Naturalis Repository

Principal publications 2001-2006 (peer reviewed papers are marked thus *):

Please contact the author when you are interested in one of the articles above and it is not available online or in the repository [Repository].