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Martien van Oijen

Dr. M.J.P. (Martien) van Oijen F.L.S., Research Associate - Taxonomy and Systematics


Email: martien.vanoijen@naturalis.nl
Phone: +31 (0)71-56 87 619
Room number: C.02.13, Darwinweg 2
“One of the things I want to do, is to unravel the history of the Naturalis collection.”

My predecessor Dr M. Boeseman used to say that it is impossible for a curator to plan anything because curators constantly have to deal with unexpected questions e.g. regarding the collection, collectors, stranded animals, odd specimens found on attics, etc. He taught me that the major tasks of a curator is to make the collection accessible to researchers, providing information about all aspects of the collection and increase the knowledge of the collection by unraveling its history. Although Naturalis did not maintain the position of curators after the separation of the sectors Research an Collections in 2005, I still feel the old responsability for the collection. Therefore I alternate pure taxonomic work (species descriptions of Lake Victoria cichlids) with work concerning the history of the collection.


After graduating as a biologist from Leiden University in June 1977, I went to Tanzania with a WOTRO grant to study haplochromine cichlids in the Mwanza Gulf of lake Victoria (East Africa). Many new species were discovered and large collections were shipped to the Rijksmusem van Natuurlijke Historie in Leiden. In October 1980, before the grant period had terminated, I obtained a permanent position as Curator of Fishes at the RMNH. As the single permanent researcher of the extensive RMNH fish collection I had extend my knowledge of fishes. In 1981 and 1983 I joined oceanographic expeditions to the Azores and the Cape Verde Islands, but a traffic accident in 1984 brought an abrupt end to my diving activities. In 1985 I joined an expedition to Lake Victoria where I witnessed the devastating impact of the Nile perch on the lake's biodiversity. In the following years I divided my time between haplochromine cichlids, Indo-pacific fishes from the Bleeker collection and Japanese fishes from the Siebold collection. In 1996 I defended my thesis on the taxonomy of piscivorous haplochromines from Lake Victoria. My knowledge of the Siebold collection resulted in a position in the Council of Experts of the SieboldHouse. I was a quest curator for three exhibitions, two of which were on fishes from the Siebold collection.  In order to make the work of Pieter Bleeker more accessible I translated with Dr G.M.P. Loots (Latin) two of his major publications. I am working on an English translation of the Fauna Japonica (Vol. Pisces).


Research interest

My training at Leiden University and the subsequent period of three years fieldwork with the Haplochromis Ecology Survey Team at the Mwanza Gulf of lake Victoria have determined my passion for fish eating and shrimp eating haplochromine cichlids. A large part of my research time is spend in making taxonomic descriptions of dozens of new (and now probably extinct) species belonging to this interesting group of fishes. The unexpected passing away of my friend and colleague Frans Witte and the impending abolition of the cichlid research at Leiden University make it probable that in the future I will more often cooperate in taxonomic research on micrognathic haplochromine species.

During my years as curator of Fishes of the Rijksmuseum van Natuurlijke historie en Naturalis (1981-2005)I developped a passion for the history of the Fish collections of this museum. Without knowledge of the history of the collectors, collecting and preservation techniques, exchange policies, and researchers who worked with the specimens it is impossible to make a good use of the collections for contempory research.  Two collections that entered  ‘s Rijks Museum van Natuurlijke Historie (the Leiden Museum) at the Rapenburg in the 19th century and were moved to new premises at the Raamsteeg in the first decades of the 20th century have my special attention.

The  most important of these is the Bleeker collection; a collection of ten thousands of fishes from the Indian archipelago assembled  between 1842 and 1860 during the period when Pieter Bleeker (1819-1878) as a medical doctor employed by the Dutch army, was (mainly) stationed in Batavia (Jakarta). Bleeker not only received fishes from colleagues and compatriots stationed at various island of the Indian archipelago, but also from further locations e.g. India, Singapore and Japan. Moreover, after his return to the Netherlands Bleeker described new species from China, Madagascar and Surinam. In more than 500 papers Pieter Bleeker described nearly 2000 new species of fishes. Unfortunately only after Bleeker passed away in 1878 the RMNH received a grant from the Dutch government to purchase the major part of Bleeker’s fish collection at the auction of the Bleeker collection iin 1879. 

Another very important collection is the collection of Japanese fishes assembled from 1823-1834 by Philipp von Siebold and Heinrich Burger.  On the basis of this collection, which also comprises 256 watercolours of fishes painted from 1830-1832 by the Japanese artist  Kawahara Keiga, the Fish Volume of the Fauna Japonica (1845) was written by the director of the RMNH C.J. Temminck and curator of vertebrates H. Schlegel. Part of the sharks and rays from this collection were described by Müller and Henle (1838-1841). Every year examination of the collection by (mostly Japanese) specialists, reveals more, previously unnoticed, species.


h. kuhl & j.c. van hasselt, east asian fishes, history of the rmnh collection, lake victoria, indo-pacific fishes, haplochromine cichlids, bleeker collection, japanese fishes, dr pieter bleeker, history of ichthyology, siebold collection

Current research topics

Taxonomic descriptions of haplochromine cichlids of Lake Victoria

When I was engaged in ieldwork in the Mwanza Gulf area Lake Victioria (1977-1980) I collected  and photographed many specimens tentatively identified as new species of piscivorous haplochrominens.  As almost all piscivores disappaered from the catches around 1985, these species are probably extinct now. Before I retire in september 2015 I want to finish the descriptions of as many of these species as possible. With a few exceptions, for instance in cases in which species are dedicated to special persons, the descriptions will be very short. 

Bleeker collection

In my last year I intend finish and publish a few papers on the hstory of the Bleeker collection, e.g on the way the fishes were preserved, and the artists who worked for  Bleeker, a paper on the types of the Japanesefish species described by bleeker, and three papers on the Chinese fishes from the Senn van basel collection described by Beeker.

Siebold collection

Research on Buerger'smanuscrips (species descriptions, shipping lists, correspondence) and Keiga paintings will soon result in a paper, and a lecture tat will be presented at the 8th International Conferennce  on the Siebold collection at Wurzburg.


  • Dr Mitsuto Aibara. Department of Biology, Faculty of Sciences, Kyushu University, Fukuoka, Japan.
  • Dr Jacco C. van Rijssel. Institute for Ecology and Evolution, University of Bern (IEE). EAWAG Center of Ecology, Evolution and Biogeochemistry (CEEB).Department of Fish Ecology and Evolution Kastanienbaum, Switzerland
  • Drs Marnix de Zeeuw Monster, The Netherlands
  • Dr P.C. Goudswaard. IMARES Wageningen UR (Instituut for Marine Resources and Ecosystem studies) Yerseke, The Netherlands
  • Dr P.A.M. van Zwieten. Aquaculture and Fisheries Group, Wageningen University, Wageningen
  • Dr Barry Russel. Museum & Art Gallery of the Northern Territory, Darwin, Australia
  • Dr Tyson Roberts. Research Associate, Smithsonian Tropical Research Institute, Panama.
  • Dr William White. CSIRO Marine & Atmospheric Research,Hobart, Australia.
  • Dr Mathias Stehmann. ICHTHYS, Ichthyological Research Laboratory and Consultant, 13, Hamburg, Germany
  • Dr Maurice Kottelat. Cornol, Swirserland
  • Dr Seishi Kimuura. Fisheries Research Laboratory, Mie University, Mie Japan
  • Dr Mari Kuroki. Department of Aquatic Bioscience, Graduate School of Agricultural and Life Sciences, The University of Tokyo
  • Dr Ryuji Hiraoka. Faculty of letters, Prefectural University of Kumamoto, Japan


In the Fish collection of the Naturalis Biodiversity Center the following collections are the most interesting for me: 

The Bleeker collection. The East Asian freshwater and marine fishes, mainly from the Indonesian archipelago, collected by Dr Pieter Bleeke from 1845 - 1860.

The Siebold collection. The Japanese freshwater and marine fishes collected by Philipp von Siebold and Heinrich Buerger from 1823 -1834.

The HEST collection The collection of haplochromine cichlids from Lake Victoria(East Africa) collected from 1977-2009



Not applicable.

Available student projects

Public outreach

News papers (in Dutch)

  • Raadselachtige haai geeft licht uit zijn achterste. door Maarten Keulemans. Volkskrant 21 april 2012.
  • Victoria's Vissen. door Ben van Raaij. Volkskrant 26 April 2014.


  • Vissen in de Oudheid 2002
  • On having no neck


Sieboldhuis 9 December 2012 Why are Japanese fishes preserved in Leiden?

Sieboldhuis 14 October 2013 Can Sushi be made of all fishes?

Science talks in LiveScience, Naturalis (2013, 2014)

Guest curator for the following exhibitions

  • Vis - stillevens van Hollanndse en Vlaamse Meesters 1550-1700. Centraal Museum 7 Feb-9 May, 2004.
  • The aquatic world of von Siebold. Nagasaki Museum of History and Culture  1 Jun- 30 juli, 2007.
  • Vissen van haai tot koi. Japanmuseum  SieblodHuis, 6 April- 8 July, 2012.



Journals SCI, peer-reviewed

Stehmann M.F.W., Oijen M. van, Kamminga P. 2016. Re-description of the rare taillight shark Euprotomicroides zantedeschia (Squaliformes, Dalatiidae), based on third and fourth record from off Chile. Cybium 40: 187-197.


Journals SCI, peer-reviewed

Tatsuya K., Oijen M.J.P. van, Kubo Y., Kitagawa E. 2015. Redescription of Ateleopus japonicus Bleeker 1853, a senior synonym of Ateleopus schlegelii van der Hoeven 1855, Ateleopus purpureus Tanaka 1915, and Ateleopus tanabensis Tanaka 1918 with designation of a lectotype for A. japonicus and A. schlegelii (Ateleopodiformes: Ateleopodidae). Zootaxa 4027: 389-407.

Kaga T., Oijen M.J.P. van, Kubo Y., Kitagawa E. 2015. Redescription of Ateleopus japonicus Bleeker 1853, a senior synonym of Ateleopus schlegelii van der Hoeven 1855, Ateleopus purpureus Tanaka 1915, and Ateleopus tanabensis Tanaka 1918 with designation of a lectotype for A-japonicus and A-schlegelii (Ateleopodiformes: Ateleopodidae). Zootaxa 4027: 389-407.
Go to website (DOI)


Journals SCI, peer-reviewed

Faria V.V., Chiaramonte G.E., Oijen M.J.P. van 2014. Publication date and author spelling for the hidden angelshark Squatina occulta. Journal of Fish Biology 84: 1185-1187.
Go to website (DOI)

Chapters in books

Kuroki M., Oijen M.J.P. van, Tsukamoto K. 2014. Eels and the Japanese: An Inseparable, Long-Standing Relationship. 91-108:


Journals SCI, peer-reviewed

Oijen M.J.P. van, Meij S.E.T. van der 2013. The types of Osteoglossum formosum Müller & Schlegel, 1840 (Teleostei, Osteoglossidae). Zootaxa 3722: 361-371.
Go to website (DOI)

Zeeuw M.P. de, Westbroek I., Oijen M.J.P. van, Witte F. 2013. Two new species of zooplanktivorous haplochromine cichlids from Lake Victoria, Tanzania. ZooKeys 256: 1-34.
Go to website (DOI)

Oijen M.J.P. van, Kawai T., Loots I. 2013. Putative type specimens of Satyrichthys (Scorpaeniformes: Peristediidae) in the Bleeker collection of the Naturalis Biodiversity Center, Leiden, The Netherlands. Zootaxa 3670: 207-214.

Naturalis Repository


Shih-Pin Huang, Martien J. P. van Oijen, Kuang-Ying Hwang and I-Shiung Chen, 2014. Redescription of Gobius gastrospilos Bleeker, 1853 with comments on four newly recorded species of brackish gobies from Taiwan. Journal of Marine Science and Technology, Vol. 21, Suppl., pp. 94-105 (2013) doi: 10.6119/JMST-013-1219-7

 V.V. Faria, G.E. Chiaramonteand M.J.P. van Oijen,  2014. Publication date and author spelling for the hidden angelshark Squatina occulta. Journal of Fish Biology (2014) 84, 1185–1187 doi:10.1111/jfb.12325, available online at wileyonlinelibrary.com


 Kuroki, Mari, Martien J.P. van Oijen & Katsumi Tsukamoto, 2013. Eels and the Japanese: An Inseparable, Long-Standing Relationship. Pp. 91-108. In Tsukamoto & Kuroki (Eds) Eels and Humans. Springer, Japan

  Oijen, Martien J.P. van & Sancia E.T. van der Meij, 2013. The types of Osteoglossum formosum Müller & Schlegel, 1840 (Teleostei, Osteoglossidae). Zootaxa 3722 (3): 361–371. http://zoobank.org/urn:lsid:zoobank.org:pub:D624BC3F-0B62-4A3F-9A6F-F3A696C71D41

 Frans Witte, Mary A. Kishe-Machumu, Olivia C. Mkumbo, Jan H. Wanink, Pleun C. Goudswaard, Jacco C. Van Rijssel, Martien J.P. van Oijen, 2013.The Fish Fauna of Lake Victoria during a Century of Human Induced Perturbationsp. 49-66 in J. Snoecks & A Getahun (Eds). Proceedings of the Fourth International Conference on African Fish and Fisheries. Zoological Online Series Royal Museum for Central Africa, Tervuren, Belgium.

 Hibino, Yusuke, Kiyotaka Hatooka, Martien J. P. van Oijen, Ronald de Ruiter, James McClaine and Seishi Kimura, 2013. Whereabouts of the worm-eel types described by Pieter Bleeker. Poster at the Symposium on systematics and diversity of Fishes, Tokyo, Japan.

Oijen, Martien van en Jacco van Rijssel. 2013. Bevlogen cichlidenexpert laat onschatbare erfenis na. In Memoriam Frans Witte. Bionieuws 2 maart 2013: 7. 

de Zeeuw M.P., Westbroek I., van Oijen M.J.P., Witte, F., 2013. Two new species of zooplanktivorous haplochrominecichlids from Lake Victoria, Tanzania. ZooKeys 256: 1–34.doi: 10.3897/zookeys.256.3871 


Oijen, M.J.P. van, & G.M.P. Loots, 2012. An illustrated translation of  Bleeker’s Fishes of the Indian Archipelago. Part II –Cyprini. Zoologische Mededelingen. 86(1): 1-469.  

Oijen, Martien J.P. van, 2012. Vissen van Haai tot Koi  pp 9-16. In: Vissen van Haai tot Koi, 16 april – 8 juli 2012. Japanmuseum SieboldHuis, Leiden. ISBN 978-90-812874-0-1

Oijen, Martien J.P. van, 2012. Fish from Shark to Koipp 26-33. In: Vissen van Haai tot Koi, 16 april – 8 juli 2012. Japanmuseum SieboldHuis, Leiden. ISBN 978-90-812874-0-1 

Hosoya Kazumi, Fujita Tomohiko, Kawase Seigo and Martien J.P. van Oijen, 2012. Riddles in the Siebold collection: Heterogeneous type series of Japanese freshwater fishes. Poster at the XIV European Congress of Ichthyology, Liège, Belgium. 


Oijen, M.J. P. van, 2011. Dagboeken uit de tijd van Philipp Franz von Siebold (4) Het hofreisverslag van J. Van Overmeer Fisscher (1822). Transcription of J. van Overmeer Fisscher (1833). Bijdrage tot de kennis van het Japanse Rijk. Daily blog at website SieboldHuis from 11 February till 7 July.  

Oijen, Martien J.P. van, Toshiyuki Suzuki and I-Shiung Chen, 2011. On the Earliest Published Species of Rhinogobius. With a Redescription of Gobius brunneus Temminck and Schlegel, 1845z. Journal of the National Taiwan Museum 64 (1):1-17.


Oijen, M.J.P. van., 2009/2010. Dagboeken uit de tijd van Philipp Franz von Siebold (3).Dagregister van het Opperhoofd Johan Wilhelm De Sturler van den 20 December 1825 tot den 5 Augustus 1826 Transcription of original diary. Daily blog at website SieboldHuis from  22 December 2009 till  6 August 2010.


Oijen, M.J.P. van & T. Raczynski, 2009. First record of a specimen of Euprotomicroides zantedeschia Hulley & Penrith, 1966, or a new species of Euprotomicroides from the South Pacific? Abstract. Book of abstracts: 8th Indo Pacific fish conference and 2009 ASFB workshop & conference 31 May - 5 June 2009 : biogeography & biodiversity. Freemantle, Western Australia Australian Society for Fish Biology; Government of Western Australia; Department of Fisheries 2009.

Oijen, M.J.P. van, 2009. Putrefied and dessicated fish specimens from the auctioned Bleeker collection in the National Natuurhitorisch Museum (RMNH) in Leiden. Abstract. Book of abstracts: 8th Indo Pacific fish conference and 2009 ASFB workshop & conference 31 May - 5 June 2009: biogeography & biodiversity. Freemantle, Western Australia Australian Society for Fish Biology ; Government of Western Australia ; Department of Fisheries 2009.

Witte, Frans, Martien J.P. van Oijen & Ferdinant A. Sibbing, 2009. Fish Fauna of the Nile. p 647-676. In: H.J.Dumont (ed.) The Nile: Origin, Environments, Limnology and Human Use. Monographiae Biologicae Vol. 89. Springer.

Oijen, M.J.P. van, 2009. Bleeker, P. A precursor of the fishes of the Indian Archipelago.Introduction. Zoologische Mededelingen. 83(1): iii-xi.

Oijen, M.J.P. van, G.M.P. Loots & F.J.G. Limburg, 2009. Bleeker, P. A precursor of the fishes of the Indian Archipelago.Translation. Zoologische Mededelingen. 83(1): 1-317.
Fransen, C.H.M.J. & M.J.P. van Oijen, 2008. L.B. Holthuis, ‘The institutional Memory’ of the Leiden Museum – Obituary. – Contributions to Zoology 77 (3): 201-204.

Oijen, M. van et alii, 2008. Herinneringen aan prof. dr. Holthuis (1921-2008). Naturalis – Nationaal Natuurhistorisch Museum 2008 (June): 1-32. [Infusis special issue.]

Oijen, M.J.P. van & M.P. de Zeeuw, 2008. Haplochromis vonlinnei spec. nov., a piscivorous haplochromine cichlid (Teleostei, Perciformes) from the Mwanza Gulf area of Lake Victoria. Zool. Med. Leiden 82 (1), 167-175.


Witte, Frans, Martien van Oijen, Mary Kishe-Machumu & Jan Wanink, 2007. Thirty Years Research on Lake Victoria Cichlids; an Ecological Drama Provides New Scientific Opportunities. – SIL news 51: 12-13.

Oijen, M.J.P. van, 2007. A short history of the Siebold collection of Japanese Fishes in the National Museum of Naural History, Leiden, The Netherlands. Catalogue of the Aquatic world of von Siebold. Nagasaki Museum of History and Culture.

Oijen, M.J.P. van, & C.S. Roselaar, 2007. Notes on types and early specimens of Bombycivora japonica von Siebold, 1824, and of Bombycilla phoenicoptera Temminck, 1828. – Zool. Meded., Leiden. 81(14),: 251-258.

Oijen, Martien J.P. van, 2007. Marinus Boeseman, 1916-2006. – Copeia 2007(1): 243-246.

Oijen, Martien J.P. van, Vicente V. Faria & Matthew T. McDavitt, 2007. The curious holotype of Pristis dubius Bleeker, 1852, and the unravelling of Bleeker’s sawfish taxonomy. Raffles Bulletin of Zoology Special edition.


Oijen, M.J.P. van & W. de Ligny, 2006 Marinus Boeseman, 1916-2006. A biography and a list of his publications. – Zool. Med. Leiden 80-4 (1):1-8, figs 1-3.

Oijen, M. J.P. van, 2006. In Memoriam Dr M. Boeseman (1916-2006). – EIS Newsletter 6: 11-12.

Chakraborty, Anirban, Martien J.P. van Oijen, Kelvin K.P. Lim & Yukio Iwatsuki, 2006.Lepturacanthus roelandti (Bleeker, 1860), a valid species of hairtail (Perciformis: Trichiuridae). Ichthyol. Res. 53: 41-46.


Oijen, M. J.P. van, 2005. Notes concerning the type specimens of Pristis zijsron Bleeker, 1851 and Pristis dubius Bleeker, 1852 and other specimens of sawfishes (Pristidae, Rajiformes) collected and described by P. Bleeker. Abstract The 7th Indo-Pacific Fish conference Abstract book. Taipei, 2005

Oijen, Martien van, Frans Witte, Kees Goudswaard & Marnix de Zeeuw, 2005. Victoriacichliden in Naturalis: in de alcohol èn in de computer. Cichlidae 31 (6): 157-163.

Oijen, Martien J.P. van, & Jos M.J. van Oijen, 2005. Von Siebold’s reis naar het hof van de shogun in het jaar 1826. Een ontdekkingstocht door Japan. Hotei publishing, Amsterdam. 254 pp.

Sudarto, Pouyaud, L. & M van Oijen, 2005. Systematics, a prerequisite for the sustainable conservation of wild populations of freshwater ornamental fish. A case study with the Asian Arowana. (Abstract)

Oijen, M.J.P. van, 2005. Data on the genesis of the Atlas Ichthyologique from a little known French paper by P. Bleeker. The Raffles Bulletin of Zoology, Supplement No. 13: 3-8.

Older publications

Oijen, M.J.P. van, 2004. Haplochromis ushindi spec. nov., the largest piscivorous cichlid in the Mwanza Gulf area of lake Victoria (East Africa) before the Nile perch upsurge. Zool. Med. Leiden 78 (1): 249-255, figs 1-4, table 1.

Motomura, H. & M.J.P. van Oijen, 2003. Polynemus kapuensis, a new threadfin (Perciformes: Polynemidae) from western Kalimantan, Indonesia, and a redescription of P multifilis Temminck & Schlegel, 1843. Zool. Med. Leiden 77 (2): 393-408.

Bor, P.H.F., M.J.P. van Oijen & C. Magenta, 2003. The egg capsule of the coral cat shark, Atelomycterus marmoratus (Bennett, 1830) (Chondrichthys: Scyliorhinidae). Zool Med. Leiden 77 (4); 87-92.

Oijen, M.J.P. van & F. Witte, 2003. A redescription of Haplochromis labriformus (Nichols & La Monte, 1938). Zool. Med. Leiden 77 (2): 473-479.

Wanink, Jan H., Edgid F.B. Katunzi, Martien J.P. van Oijen & Frans Witte, 2002. Adaptive responses of fish species surviving arnthropogenic perturbation in Lake Victoria. – Abstracts for Aquatic Ecology.

Oijen, M.J.P. van, 2001. Squalus edwardsii (currently Haploblepharus edwardsii: Chondrichthyes, Carcharhiniformes): proposed attribution to Schinz (1822) and conservation of edwardsii as the correct spelling of the specific name. Bull. Zool. Nom. 58 (4): 294-296.

Motomura, H., M.J.P. van Oijen, I.J.H. Isbrücker & Y. Iwatsuki, 2001. Redescription of a rare threadfin (Perciformes: Polynemidae), Polydactylus macrophthalmus (Bleeker, 1858), with designation of a lectotype and notes on distributional implications. Ichthyol. Res. 48: 289-294.

Oijen, M.J.P. van, 2000. Badis badis (Hamilton, 1822), een bron van misverstanden. - Het Aquarium 70 (20): 71-73.

Van den Hoek Ostende, L., C. Smeenk, R. Dekker, M. Hoogmoed & M. van Oijen, 1999. Swift as a shadow. Houghton Mifflin Company, New York. 158 pp.

Oijen, M.J.P. van, 1999. On the proposed designation of a single neotype for Hemibagrus nemurus (Valenciennes, 1840) (Osteichthyes, Siluriformes) and H. sieboldii (Bleeker, 1846), and of the lectotype of H. planiceps (Valenciennes, 1840) as a neotype for H. flavus (Bleeker, 1846). Bull. Zool. Nom. 56 (3): 200-201.

Witte, F., Barel, C.D.N. & M.J.P. van Oijen, 1997. Intraspecific variation of haplochromine cichlids from Lake Victoria and its taxonomic implications. South African Journal of science 93: 585-593.

Oijen, M.J.P. van, & F. Witte, 1996. Taxonomical and ecological description of a species complex of zooplanktivorous and insectivorous cichlids from Lake Victoria.-- Zool. Verh. Leiden 302: 1-56.

Oijen, M.J.P. van, 1996. The generic classification of the haplochromine cichlids of Lake Victoria, East Africa.-- Zool. Verh. Leiden 302: 57-110.

Witte, F. & M.J.P. van Oijen, 1995. Biology of haplochromine trophic groups: 321-335. In: F. Witte & W.L.T. van Densen (eds). Fish stocks and fisheries of Lake Victoria. A handbook for field observations.-- Cardigan, Dyfed, U.K.

Oijen, M.J.P. van, 1995. Key to Lake Victoria fishes other than haplochromine cichlids: 209-298. In: F. Witte & W.L.T. van Densen (eds). Fish stocks and fisheries of Lake Victoria. A handbook for field observations-- Cardigan, Dyfed, U.K.

Crul, R.C.M., G.T. Silvester, D.J. Postma, M.J.P. van Oijen, T.O. Acere & G. Bongers, 1995. A bibliography of Lake Victoria. i-viii, 1-164.-- UNESCO, Paris.

Witte, F., T. Goldschmidt, P.C. Goudswaard, W. Ligtvoet, M.J.P. van Oijen & J.H. Wanink, 1992. Species extinction and concomitant ecological changes in Lake Victoria. Neth. J. zool.42 (2-3): 214-232.

Witte, F., T. Goldschmidt, J. Wanink, M. van Oijen, K. Goudswaard, E. Witte-Maas& N. Bouton,1992. The destruction of an endemic species flock: quantitative data on the decline of the haplochromine cichlids of Lake Victoria. Env. Biol. Fish. 34:1-28.

Oijen, M.J.P. van, 1992. Haplochromis howesi spec. nov., a crab and fish eating cichlid from lake Victoria. Zoologische Mededelingen Leiden 66 (40): 561-579.

Oijen, M.J.P. van, 1992. Het Victoriameer: van cichlidenoase tot Nijlbaarswoestijn. –Aquariumwereld 45 (1): 3-6.

Oijen, M.J.P. van, 1991. A systematic revision of the piscivorous haplochromine cichlidae ( Pisces: Teleostei) of Lake victoria (East Africa). Part 1. Zool. Verh. Leiden 272: 1-95.Check-List of the Freswater Fishes of Africa. ORSTOM, Paris.Check-List of the Freswater Fishes of Africa. ORSTOM, Paris.Check-List of the Freswater Fishes of Africa. ORSTOM, Paris.Check-List of the Freswater Fishes of Africa. ORSTOM, Paris.Check-List of the Freshwater Fishes of Africa. ORSTOM, Paris.

Oijen, M.J.P. van, J. Snoeks, P.H. Skelton, C. Maréchal & G.G.Teugels, 1991. Haplochromis (pp. 100-184). In: Daget, J. J.P. Gosse, G.G. Teugels & D.F.E. Thys van den Audenaerde (eds). Check-List of the Freshwater Fishes of Africa. ORSTOM, Paris.

Oijen, M.J.P. van, 1991. On the high percentage of piscivorous cichlids of lake Victoria. Abstract. – Bull. Zool. Mus. Special issue August 1991: 54.

Witte, F. & M.J.P. van Oijen, 1990. Taxonomy, ecology and fishery of Lake Victoria haplochrimine trophic groups. – Zool. Verh. Leiden262:1-47

Oijen, M.J.P. van, 1990. Het Victoriameer: van cichlidenoase tot Nijlbaarswoestijn. – Panda 26 (4): 3-5.

Oijen, M.J.P. van, 1989. Notes on the relationship between prey and pharyngeal jaw size in piscivorous haplochromine cichlids from Lake Victoria. – Ann. Kon. mus. Mid. Afr., Zool. Wetensch. 257: 73-76.

Wheeler, A. & M.J.P. van Oijen, 1985 The occurrence of Sphoeroides pachygaster (Osteichthyes- Tetraodontiformes) off North-West Ireland. Zool. Med. Leiden 59 (11): 101-107.

Oijen, M.J.P. van, 1983. Het Victoriameer; meer dan mooi om naar te kijken. – N.V.C. periodiek 9(5): 1-14.

Oijen, M.J.P. van, 1982. Ecological differentiation among the piscivorous haplochromine cichlids of Lake Victoria. Neth. J. Zool. 32: 336-363.

Oijen, M.J.P. van, F. Witte & E.L.M. Witte-Maas, 1981. An introduction to ecological and taxonomic investigations on the haplochromine cichlids from the Mwanza Gulf of lake Victoria. Neth. J. Zool. 31: 149-174.

Oijen, M.J.P. van, 1978. The live colouration of certain previously described Haplochromis species: 180-192. In P.H. Greenwood, & C.D.N. Barel. A revision of the Lake Victoria Haplochromis species (Pisces, Cichlidae) Pt III . Bull. Br. Mus. (Nat. Hist.) Zool. 33 (2).

Barel, C.D.N., M.J.P. van Oijen, F. Witte & E.L.M. Witte-Maas, 1977. An introduction to the taxonomy and morphology of the haplochromine cichlids from Lake Victoria. Neth. J. Zool. 27 (4): 333-389.

Barel, C.D.N., F. Witte & M.J.P. van Oijen, 1976. The shape of the skeletal elements in the head of a generalized Haplochromis species: H. elegans Trewavas 1933 (Pisces, Cichlidae). Neth. J. Zool. 26 (2): 163-265.

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