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Lars van den Hoek Ostende

Dr. L.W. (Lars) van den Hoek Ostende, Researcher - Biodiversity Discovery, Fossil microvertebrates

Contact

Email: lars.vandenhoekostende@naturalis.nl
Phone: +3171 751 9685
Room number: Vondellaan 55
“Whereas policy makers see competition as a tool for excellence, true progress in science can only be made by cooperation”

As most vertebrate palaeontologists, my fascination for lost fossils must have started with dinosaurs. I outgrew that phase. In order to understand the history of our planet, you cannot focus on one, exceptional fossil; you need many. Micromammals are ideally suited for recording changes in the biosphere, particularly in the period which holds my greatest interest, the Neogene. In trying to understand the development of (mammal) biodiversity of that time, my research revolves around the question "What was when where and why". Admittedly, that is a rather broad question. As a trained taxonomist, with a peculiar appetite for fossil insectivores, I was traditionally focussed on the what. Over the years, I have connected and interconnected  with many colleagues in the field in order to gain the wider scope and to focus on the question that really matter: Why? I am privileged in being allowed to work with so many fine colleagues and young talents from all over the world.

Career

Science is a state of mind, not a profession. If that is true, I am a scientist working at Naturalis for over 25 years. My resume tells differently, since I only have been employed as a scientist at the institute since 2002. During that period I also had my share of management, as a departmental head for over two years. Before my appointment as a researcher, I was a jack-of-all-trades for a decade, serving contracts including educational duties, collection management, editorial work, and whatever needed to be done. It was during those days that I wrote my PhD in the off-hours. Science is a state of mind.

Research

Research interest

My research revolves around mammal biogeography of the Miocene and the ecosystem dynamics of that period. In addition, I dabble a bit in the insular evolution of micromammals.

Keywords

stratigraphy, miocene, insectivores, neogene, insular evolution, palaeobiogeography, palaeoclimatology

Current research topics

The Miocene of Europa and Anatolia

A couple of years ago, I returned to the region on which I did my PhD. Anatolia, strategically positioned between Asia, Europe and Africa, is an incredibly interesting area from a biogeographical point of view. The more so in the early Miocene, when vast changes took place in the topography of the region, including the closure of the eastern Tethys, leading to the formation of the Mediterranean. I had the great pleasure of leading a National Geographic expedition into the Anatolian Turkish Miocene and am currently working on the results of these and other excavations with my partners Serdar Maya (EGE University, Izmir), Pablo Pelaez Campomanes (Museo Nacional de Ciencas Naturales, Madrid), Peter Joniak (Comenius University, Bratislava) and other talented co-workers.

Our work in the eastern Mediterranean is particularly of interest in combination with work done elsewhere in Europe. I have been, and continue to be, active in the Iberian Peninsula with colleagues from Madrid and Barcelona. Covering both the east and the west of the Mediterranean allows us to reconstruct change in the environment through space and time. There are many questions concerning provincialism, faunal turnovers and evolutionary scenarios. For me personally, the most tantalizing is why there is such a big temporal gap between the largest topographical change (i.e., the Africa/Eurasia collision) and the major climate event (i.e. the mid-Miocene cooling).

 

A sea of data

Modern science is faced with the tremendous challenge of extracting patterns out of an even increasing sea of data. Particularly when dealing with a complex topic as ecosystem evolution, this is a central point of concern and a source of opportunities at the same time. As coordinator of the RCMNS working group Terrestrial Ecosystems, I aim to promote cooperation between research groups to combine data from various sources. It is my ambition to enable studies that will reconstruct palaeoenvironments based on multiple proxies and to merge such studies into a comprehensive view of the changes in the western Eurasian Neogene ecosystems.

Continuous progress in numerical methods provide us with the means to study changes in the biosphere in ever increasing detail, using large datasets. Whereas I very much applaud this development, it also raises some concern. In pursuit of innovations, scientist more and more need to rely on data, while the basic study of fossils, i.e. the data gathering, receives less and less attention. Since a old-fashioned analysis on a good dataset is more likely to provide reasonable result than an innovative method on a poor one, preservation of data-quality is very much a point of focus, both in my teaching and in my scientific work. And this goes double for my role as assistant coordinator for the New and Old World database. Hence, much of my studies still deal with the basic taxonomy of the group I am most familiar with, the insectivores (Eulipotyphla).

 Desmanella gudrunae

 

Insular evolution 

Dutch palaeontologists have a long tradition in the study of island faunas. For instance, Naturalis houses the largest collection of fossils from the Neogene islands of Gargano (Italy). As many fellow scientists, I am convince that the driving force behind insular evolution are adaptations pertaining to a change in life strategy. At the same time I am acutely aware that the focus on insular evolution has been always on the fascinating examples, whereas the process in its entirety can only be understood if we also take into account the species that seem to have been less or even un-affected by the island rule. This line of research is at present mainly maintained through the work of students.

 A fragment of the lower jaw of Papagomys spec., a giant rat from the island of Flores (Indonesia)

And all that jazz

As I mentioned before, I started my career as a Jack-of-all-trades. In retrospect, I was fortunate with this ‘false start’, as it kindled many interests very much alive today. And I learned it fits my personality to be also actively involved in teaching, popular science, editing and all kinds of fun stuff. Multitasking is in the blood and where I explain my research ‘focus’ above, don’t be surprised to find the occasional paper on Caribbean fossil bats or some amazing fossils from Siberia. Entering the final stages of the career, the mind seems to wander a bit more and the overall picture more relevant than the details. Those are better left to a younger generation, to which I am now a teacher and mentor. Quality needs to be preserved. As such, I am nowadays more focussed on sharing my experience, and I do have some stories to tell, some dreams to share and some roads to explore with good companions.

Collaborations

Currently I am involved in projects with the Museo Nacional de Ciencias Naturales/CSIC (Madrid), EGE University (Izmir), Institut Catalá de Paleontologiá (Barcelona) and Comenius University (Bratislava). But the list of people I had the immense pleasure of working with is much longer. Science if fun, and fun needs to be shared. Or maybe it is the sharing that makes it fun. And for the collaboration over the last decades I certainly need to thank Pablo, Marc, Serdar, Isaac, Peter, Maya, Marian, Doris, Vero, Adri, Paloma, Jerome, Daniela, Daniela, Delia, Rutger, Steven, Anneke, Francien, David, Elisa, Melike, Janos, Constantin, Jorge,  Johannes, to name but a few of my favorite people.

Supervision PhDs

Co-promotor/supervisor

I have co-supervised the PhD of Elisa (Ferrera). Currently I am supervising the PhD of Anneke Madern (Leiden) and co-supervising the PhD of Daniela Winkler (Hamburg)

Teaching

Courses

Since 2006, I have been coordinating the palaeontology teaching in BSc and MSc courses at Leiden University, and teaching part of the curriculum myself.

Supervision MSc Students

2017 Supervision Gosse de Vries
2016 Supervision Jacqueline Schroeder
2016 Supervision Renda Remmerswaal
2015 Supervision Jorn Bom
2013 Supervisor - Jeanne van de Put - Vallesian micromammal biogeography of the Iberian Peninsula. Comparing biodiversity through time and space between the Vallès-Penedès basin and inland basins.
2013 Supervisor Flora van Glabbeek - The phylogeny of Erinaceidae: Obtaining a complete picture by a combination of morphological and DNA methodologies.
2013 Supervisor Jesse Hennekam - Dental variation in the island ruminant Hoplitomeryx
2013 Supervisor Lussi Jordanov - Mesowear reconstruction of Candiacervus.
2013 Supervisor Lysanne van Bennekom - The micromammals of Eriçek (Pliocene, Anatolia)
2013 Supervisor Pavlos Piskoulis - Revision of the Late Pleistocene/Holocene micromammal fauna from Marie-Jeanne Cave (Hastière-Lavaux, Belgium)
2012 Supervision Leonie Bergwerff- The effects of the Vallesian Crisis on the Carnivora of the Vallès-Penedès Basin.
2012 Supervision Naomi Janssen - The effects of the Vallesian Crisis on the Artiodactyla of the Vallès-Penedès Basin.
2012 Supervisor Flemming Diepeveen - The micromammals from Biçakçi (Anatolia).
2012 Supervision Suzanne van der Hoeven - Revision of the vole Kislangia rex.
2010 Supervisor Anne Blair Gould - Premature death of happy old age? Reconstructing the life histories of extinct rodents
2010 Supervisor David Schuuring - Palaeontological techniques, general knowledge (Archaeology internsip)
2009 Supervisor Anneke Madern - Biodiversity patterns and hotspots of Miocene mammals
2008 Supervisor Roeli Zandvliet - Micromammals of Song Gupuh, Java
2006 Supervisor Wishal Baktawar - Flying foxes (Chiroptera, Mammalia) from Liang Bua
2005 Supervisor Esther van der Wel - The postcranial skeleton of Deinogalerix

Supervision BSc Students

2017 Supervision Rianne van Stappen
2015 Supervisor - Sanne ten Thije - Accuracy of MN zonation type locality system
2015 Supervision Renda Remmerswaal
2015 Supervision Magali Bouquet
2015 Supervisor Guido Smit
2012 Supervisor Flora van Glabbeek - Morphological differences in the dentition fo the Dimylidae
2010 Supervisor Mariska Koekoek - Small mammals of Gargano (Italy) from the Late Miocene; do they show change in size.
2010 Supervisor Linda Gilein - Pleistocene voles of Beremend 15
2010 Supervisor Rogier van Oostrum - aDNA extraction and mtDNA sequencing of endemic Murinae from the Liang Bua Cave, Flores, Indonesia
2009 Supervisor Marlous Ouwendijk - The microchiroptera of Liang Bua (Flores, Indonesia)
2007 Supervisor Arike Gill - An overview of fossil rodents from Aruba
2007 Supervisor Anneke Madern - Fossil Pleistocene rodent fauna from Aruba and Bonaire
Available student projects

Public outreach

The greatest assett of our species is our curiosity. Within our society, scientists play to role of satisfying that curiosity, and therefore sharing their knowledge with society is part of the work package of any scientist. Moreover, every scientist likes to talk about his work.

In my early career I wrote popular articles for several magazines, and was involved in a number of exhibitions. More and more, my popular writing took the form of columns, of which I have produced many. Unfortunately, this part of the work has over the last couple of year fallen victim to the concept of performance parameters. So mostly my outreach activities are now limited to interviews, being in the fortunate position that the worlds of the past tickle the curiosity of the general public. Still, the road to the future is paved with good intentions, and I sure hope to pick up popular writing again.

Publications

2018

Journals SCI, peer-reviewed

Alçiçek M.C., Hoek Ostende L.W. van den, Saraç G., Tesakov A.S., Murray A.M., Hakyemez H.Y., Goktas F., Mayda S., Jimenez-Moreno G., Buyukmeric Y., Wesselingh F.P., Alçiçek H. 2018. Comment on "Miocene to Quaternary tectonostratigraphic evolution of the middle section of the Burdur-Fethiye Shear Zone, south-western Turkey: Implications for the wide inter-plate shear zones. Tectonophysics 690, 336-354". Tectonophysics 722: 595-600.

Madern P.A., Put J.M.M.S. van der, Casanovas-Vilar I., Hoek Ostende L.W. van den 2018. Iberian micromammals show local extent of Vallesian Crisis. Palaeogeography, Palaeoclimatology, Palaeoecology 496: 18-31.

Jovells-Vaqué S., García-Paredes I., Furió M., Angelone C., Hoek Ostende L.W. van den, Berrocal Barberà M., DeMiquel D., Madurell-Malapeira J., Casanovas-Vilar I. 2018. Les Cases de la Valenciana, a new early Miocene small-mammal locality from the Vallès-Penedès Basin (Catalonia, Spain). Historical Biology 30: 404-421.
Go to website (DOI)

Villa A., Blain H.-A., Hoek Ostende L.W. van den, Delfino M. 2018. Fossil amphibians and reptiles from Tegelen (Province of Limburg) and the early Pleistocene palaeoclimate of The Netherlands. Quaternary Science Reviews 187: 203-219.

Journals non-SCI, peer-reviewed

Furió M., Hoek Ostende L.W. van den, Minwer-Barakat R. 2018. Evolución de las asociaciones de insectívoros (Eulipotyphla, Mammalia) en España y su relación con los cambios climáticos del Neógeno y el Cuaternario.. 27: 38-51.
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2017

Journals SCI, peer-reviewed

Alçiçek H., Wesselingh F.P., Alçiçek M.C., Jimenez-Moreno G., Feijen F.J., Hoek Ostende L.W. van den, Mayda S., Tesakov A.S. 2017. A multiproxy study of the early Pleistocene palaeoenvironmental and palaeoclimatic conditions of an anastomosed fluvial sequence from the Çameli Basin (SW Anatolia, Turkey). Palaeogeography, Palaeoclimatology, Palaeoecology 467: 232-252.
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Hír J., Vencel, M., Codrea V., Rössner G.E., Angelone C., Hoek Ostende L.W. van den, Rosina V.V., Kirscher U., Prieto J. 2017. Badenian and Sarmatian s. str. from the Carpathian area: Taxonomical notes concerning the Hungarian and Romanian small vertebrates and report on the ruminants from the Felsotarkany Basin. Comptes Rendus Palevol 16: 312-332.

Hoek Ostende L.W. van den, Alvarez-Sierra M.A., García-Paredes I., Montoya P., Ruiz-Sanchez F.J., Pelaez-Campomanes P. 2017. Alto de Ballester, biogeographical consequences of atypical MN 3 micromammal assemblages from eastern Spain. Palaeontographica Abt. A 308: 127-175.

Hoek Ostende L.W. van den, Geer A.A.E. van der, Wijngaarden C.L. 2017. Why are there no giants at the dwarves feet? Insular micromammals in the eastern Mediterranean. Quaternary International 445: 269-278.

2016

Journals SCI, peer-reviewed

Jiménez-Moreno G., Alçiçek H., Alçiçek M.C., Hoek Ostende L. van den, Wesselingh F.P. 2016. Corrigendum to "Vegetation and climate changes during the late Pliocene and early Pleistocene in SW Anatolia" (vol 84, pg 448, 2015). Quaternary Research 85: 185-186.
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Hoek Ostende L.W. van den, Furió M., Madern A., Prieto J. 2016. Enters the shrew, some considerations on the Miocene palaeobiogeography of Iberian insectivores. Comptes Rendus Palevol 15: 813-823.
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Jiménez-Moreno G., Alçiçek H., Alçiçek M.C., Hoek Ostende L. van den, Wesselingh F.P. 2016. Vegetation and climate changes during the late Pliocene and early Pleistocene in SW Turkey–Response to comments by Elitez et al., Quaternary Research, 84, pp. 448–456. Quaternary Research 85: 476-477.
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Casanovas-Vilar I., Flynn L.J., Hoek Ostende L.W. van den 2016. Windows into deep time – Cenozoic faunal change in long continental records of Eurasia. Comptes Rendus Palevol 15: 753-762.

García-Paredes I., Alvarez-Sierra M.A., Hoek Ostende L.W. van den, Hernandez-Ballerin V., Hordijk K., López-Guerrero P., Oliver A., Pelaez-Campomanex P. 2016. The Aragonian and Vallesian high-resolution micromammal succession from the Calatayud-Montalbán Basin (Aragón, Spain). Comptes Rendus Palevol 15: 781-789.

Casanovas-Vilar I., Madern P.A., Alba D.M. Cabrera L., García-Paredes I., Hoek Ostende L.W. van den, DeMiguel D., RRobles J.M., Furió M., Dam J. van, Garcés M., Angelone C., Moyà-Solà S. 2016. The Miocene mammal record of the Vallès-Penedès Basin (Catalonia). Comptes Rendus Palevol 15: 791-812.
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Hír J., Venczel M., Codrea V., Angelone C., Hoek Ostende L.W. van den, Kirschner U., Prieto J. 2016. Badenian and Sarmatian s.str. from the Carpathian area: Overview and ongoing research on Hungarian and Romanian small vertebrate evolution. Comptes Rendus Palevol 15: 863-875.
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Villa A., Rozek Z., Tschopp E., Hoek Ostende L.W. van den, Delfino M. 2016. Palaeobatrachus euridyces, sp. nov. (Amphibia, Anura), last western European palaeobatrachid.. Journal of Vertebrate Paleontology 36: e1211664.
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2015

Journals SCI, peer-reviewed

Madern P.A., Hoek Ostende L.W. van den 2015. Going south: Latitudinal change in mammalian biodiversity in Miocene Eurasia. Palaeogeography, Palaeoclimatology, Palaeoecology 424: 123-131.
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Klietmann J., Hoek Ostende L.W. van den, Nagel D., Rummel M. 2015. Insectivore palaeoecology: a case study of a Miocene fissure filling in Germany. Palaeogeography, Palaeoclimatology, Palaeoecology 418: 278-289.
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Furió M., Prieto J., Hoek Ostende L.W. van den 2015. Three million years of "Terror-Shrew" (Dinosorex, Eulipotyphla, Mammalia) in the Miocene of the Valles-Penedes Basin (Barcelona, Spain). Comptes Rendus Palevol 14: 11-124.
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Alcalde G.M., Hoek Ostende L.W. van den 2015. Above the hoof: functional morphology of the fossil deer from Tegelen, the Netherlands. Geological Journal 50: 271-289.
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Hoek Ostende L.W. van den, Diepeveen F., Tesakov A., Sarac G., Mayhew D., Alcicek M.C. 2015. On the brink: micromammals from the latest Villanyian from Bicakci (Anatolia). Geological Journal 50: 230-245.
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Hoek Ostende L.W. van den, Mayda S., Oliver A., Madern A., Hernández-Ballarín V., Peláez-Campomanes P. 2015. Aliveri revisited, a biogeographical appraisal of the early Miocene mammals from the eastern Mediterranean. Palaeobiodiversity and Palaeoenvironments 95: 271-284.
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Hoek Ostende, L.W. van den, Fejfar, O. 2015. All time high: Dimylidae (Eulipotyphla, Mammalia) diversity in the early Miocene locality of Ahníkov 1 (Czech Republic,MN3). Palaeobiodiversity and Palaeoenvironments 95: 453-464.

Hordijk K., Bosma A., Bruijn H. de, Dam J. van, Geraedts C., Hoek Ostende L., Reumer J., Wessels W. 2015. Biostratigraphical and palaeoecological implications of the small mammal assemblage from the late early Miocene of Montalvos2, Teruel Basin, Spain. Palaeobiodiversity and Palaeoenvironments 95: 321-346.

Locatelli E., Due R.A., Jatmiko Hoek Ostende L.W. van den 2015. Middle-sized murids from Liang Bua (Flores, Indonesia): insular endemics, human introductions and palaeoenvironment. Palaeobiodiversity and Palaeoenvironments 95: 497-512.

Prieto J., Hoek Ostende L.W. van den, Hír J., Kordos, L. 2015. The Middle Miocene insectivores from Hasznos (Hungary, Nógrád County). Palaeobiodiversity and Palaeoenvironments 95: 431-451.

Hoek Ostende L.W. van den, Gardner J.D., Bennekom L. van, Alçiçek M.C., Murray A., Wesselingh F.P., Alçiçek H., Tesakov A. 2015. Ericek, a new Pliocene vertebrate locality in the Çameli Basin (southwestern Anatolia, Turkey). Palaeobiodiversity and Palaeoenvironments 95: 305-320.

Hoek Ostende L.W. van den, Peláez Campomanes P., Wessels W. 2015. Introduction to the special issue "Old worlds, new ideas. A tribute to Albert van der Meulen". Palaeobiodiversity and Palaeoenvironments 95: 247-252.

Kolb C., Scheyer T.M., Veitschegger K., Forasiepi A.M., Amson E., Van der Geer A.A.E., Van den Hoek Ostende L.W., Hayashi S., Sanchez-Villagra M.R. 2015. Mammalian bone palaeohistology: a survey and new data with emphasis on island forms. PeerJ 3: e1358.
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Klietmann J., Nagel D., Rummel M., Van den Hoek Ostende L.W. 2015. A gap in digging: the Talpidae of Petersbuch 28 (Germany, early Miocene). Paläontologische Zeitschrift 89: 563-592.

Jiménez-Moreno G., Alçiçek H., Alçiçek M.C., Hoek Ostende L. van den, Wesselingh F.P. 2015. Vegetation and climate changes during the late Pliocene and early Pleistocene in SW Anatolia, Turkey. Quaternary Research 84: 448–456.
Go to website (DOI)

2014

Journals SCI, peer-reviewed

Prieto J., Angelone C., Casanovas-Vilar I., Gross M., Hir J., Hoek Ostende Lars W. van den, Maul Lutz C., Vasilyan D. 2014. The small mammals from Gratkorn: an overview. Palaeobiodiversity and Palaeoenvironments 94: 135-162.
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Casanovas-Vilar I., Hoek Ostende L.W. van den, Furio M., Madern P.A. 2014. The range and extent of the Vallesian Crisis (Late Miocene): new prospects based on the micromammal record from the Valles-Penedes basin (Catalonia, Spain). Journal of Iberian Geology 40: 29-48.
Go to website (URI)

Klietmann J., Nagel D., Rummel M. Hoek Ostende L.W. van den 2014. Heterosorex and Soricidae (Eulipotyphla, Mammalia) of the fissure Petersbuch 28; micro-evolution as indicator of temporal mixing?. Comptes Rendus Palevol 13: 157-181.
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Klietmann J., Nagel D., Rummel M., Hoek Ostende L.W. van den 2014. Amphiperatherium and Erinaceidae of Petersbuch 28. Bulletin of Geosciences 89: 1-20.
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Geer A.A.E. van der, Lyras G.A., Hoek Ostende L.W. van den, Vos J. de, Drinia H. 2014. A dwarf elephant and a rock mouse on Naxos (Cyclades, Greece) with a revision of the palaeozoogeography of the Cycladic Islands (Greece) during the Pleistocene. Palaeogeography, Palaeoclimatology, Palaeoecology 404: 133-144.
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Ouwendijk M., Hoek Ostende L.W. van den, Donovan S.K. 2014. Fossil bats from the Late Pleistocene Red Hills Road Cave, Jamaica. Caribbean Journal of Science 47: 284-290.

Klietmann J., Nagel D., Rummel M., Hoek Ostende L.W. van den 2014. Enlightning complexity. The Dimylidae of Petersbuch 28. Palaeobiodiversity and Palaeoenvironments 94: 463-479.
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Mayhew D.F., Dieleman F.E., Slupik A.A., Hoek Ostende L.W. van den, Reumer J.W.F. 2014. Small mammal assemblages from the Quaternary succession at Moriaanshoofd (Zeeland, the Netherlands) and their significance for correlating the Oosterschelde fauna. Netherlands Journal of Geosciences - Geologie en Mijnbouw 93: 119-134.
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Geer A.A.E. van der, Lyras G.A., Hoek Ostende L.W. van den, Vos J. de, Drinia H. 2014. A dwarf elephant and a rock mouse on Naxos (Cyclades, Greece) with a revision of the palaeozoogeography of the Cycladic Islands during the Pleistocene. Palaeogeography, Palaeoclimatology, Palaeoecology 404: 133-144.
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2013

Journals SCI, peer-reviewed

Villier B., Hoek Ostende L.W. van den, Vos J. de, Pavia M. 2013. New discoveries on the giant hedgehog Deinogalerix from the Miocene of Gargano (Apulia, Italy). Geobios 46: 63-75.
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Freudenthal M., Hoek Ostende L.W. van den, Martin-Suarez E. 2013. When and how did the Mikrotia fauna reach Gargano (Apulia, Italy)?. Geobios 46: 105-109.
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Donovan S.K., Hoek Ostende L.W. van den 2013. Roamerella, a new name for the genus Roemerella Akopyan in Akopyan et al., 1990 (Mollusca, Gastropoda; Upper Cretaceous), preoccupied by Roemerella Hall and Clarke, 1890 (Brachiopoda, Lingulata; Devonian). Cretaceous Research 40: 1-2.
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Klietmann J., Metscher B.D., Hoek Ostende L.W. van den, Nagel D., Rummel M. 2013. First record of an upper deciduous molar in Desmanella (Uropsilinae, Talpidae, Mammalia). Geobios 46: 503-510.

Winkler D., Hoek Ostende L.W. van den, Schultz E. 2013. Dietary divergence in space and time – Lessons from the dwarf-goat Myotragus balearicus (Pleisto-Holocene, Mallorca, Spain).. Mammalian Biology 78: 430-437.

Klietmann J., Nagel D., Rummel M., Hoek Ostende L.W. van den 2013. Tiny teeth of consequence: vestigal antemolars provide key to Early Miocene soricid taxonomy (Eulipotyphla, Soricidae). Comptes Rendus Palevol 12: 257-267.

Madern A., Casanovas-Vilar I., Alba D.M., DeMiguel D., Robles J.M., Hoek Ostende L.W. van den, Moyà-Solà S. 2013. The abrupt collapse of a diversity hotspot? Reconsidering Vallesian (Late Miocene) diversity in its type area. Journal of Vertebrate Paleontology 33: 168.

Journals non-SCI, peer-reviewed

Donovan S.K., Baalbergen E., Ouwendijk M., Paul C.R.C., Hoek Ostende L.W. van den. 2013. Review and prospectus of the Late Pleistocene fauna of the Red Hills Road Cave, Jamaica. Cave and Karst Science 40: 79-86.

Alvarez-Sierra M.A., Garcia-Paredes I., Hernandez-Ballerin V., Hoek Ostende L.W. van den, Hordijk K., Lopez-Guerrero P., Meulen A.J. van der, Oliver A., Pelaez-Campomanes P. 2013. Models of historical biogeography and continental biochronology. Spanish Journal of Palaeontology 28: 129-138.

2012

Journals SCI, peer-reviewed

Ouden N. den, Reumer J.W.F., Hoek Ostende L. W. van den 2012. Did mammoth end up a lilliput? Temporal body size trends in Late Pleistocene Mammoths, Mammuthus primigenius (Blumenbach, 1799) inferred from dental data. Quaternary International 255: 53-58.
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Hoek Ostende L. W. van den, Lopez-Guerrero P., Pelaez-Campomanes P., Alvarez-Sierra M.A., Garcia-Paredes I. 2012. Early Late Miocene insectivores (Eulipotyphla, Mammalia) from the Canada section (Province of Zaragoza, east Central Spain). Comptes Rendus Palevol 11: 495-506.
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Prieto J., Hoek Ostende L.W. van den, Hir J. 2012. The Middle Miocene insectivores from Samsonhaza 3 (Hungary, Nograd County): Biostratigraphical and palaeoenvironmental notes near to the Middle Miocene Cooling. Bulletin of Geosciences 87: 227-240.
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Meulen A. J. van der, Garcia-Paredes I., Alvarez-Sierra M. A., Hoek Ostende L. W. van den, Hordijk K., Oliver A., Pelaez-Campomanes P. 2012. Updated Aragonian biostratigraphy: Small Mammal distribution and its implications for the Miocene European Chronology. Geologica Acta 10: 1-24.
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Donovan S.K., Hoek Ostende L.W. van den 2012. Neoextrania, a replacement name for Extrania Sun & Wang, 1985 (Ostracoda), preoccupied by Extrania Qian in Qiu et al., 1983 (Trilobita). Alcheringa: An Australasian Journal of Palaeontology 36: 423-423.
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Locatelli E., Due R.A., Bergh G.D van den, Hoek Ostende, L.W. van den 2012. Pleistocene survivors and Holocene extinctions: The giant rats from Liang Bua (Flores, Indonesia). Quaternary International 281: 47-57.
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Meijer H.J.M., Gill A., Louw P.G. de ,Hoek Ostende L.W. van den, Hume J.P., Rijsdijk K.F. 2012. Dodo remains from an in situ context from Mare aux Songes, Mauritius. Die Naturwissenschaften 99: 177-184.
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Casanovas-Vilar I., Hoek Ostende L.W. van den, Furió M., Madern P.A. 2012. Patterns as pretty as can be: the range and extent of the Vallesian Crisis (Late Miocene) in the Vallès-Penedès basin (Catalonia, Spain). Journal of Vertebrate Paleontology 32: 75-75.

Journals non-SCI, peer-reviewed

Bruijn H. de, Doukas C.S., Hoek Ostende L.W. van den, Zachariasse J.W. 2012. New finds from rodents and insectivores from the Upper Miocene at Plakias (Crete, Greece). Schweizerische Paläontologische Abhandlungen 131: 61-75.

2011

Journals SCI, peer-reviewed

Meulen A.J. van der, Garcia-Paredes I., Angeles Alvarez-Sierra M., Hoek Ostende L.W. van den, Hordijk K., Oliver A., Lopez-Guerrero P., Hernandez-Ballarin V., Pelaez-Campomanes P. 2011. Biostratigraphy or biochronology? Lessons from the Early and Middle Miocene small Mammal Events in Europe. Geobios 44: 309-321.
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Furió M., Casanovas-Vilar I., Hoek Ostende L.W. van den 2011. Predictable structure of Miocene insectivore (Lipotyphla) faunas in Western Europe along a latitudinal gradient. Palaeogeography, Palaeoclimatology, Palaeoecology 304: 219-229.
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Hoek Ostende L. W. van den, Reumer J.W.F. 2011. Hylopetes magistri, new name for Hylopetes debruijni Reumer & Van den Hoek Ostende, 2003, preoccupied. Journal of Vertebrate Paleontology 31: 928-928.
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Lopez-Guerrero P., Garcia-Paredes I., Hoek Ostende L.W. van den, Dam J.A. van, Alvarez-Sierra M.A., Hernandez-Ballarin V., Meulen A.J. van der, Oliver A., Pelaez-Campomanes P. 2011. Canada: a new micromammal succession from the lower Vallesian and Turolian of the Daroca area (Calatayud-Montalban basin, Spain). Estudios Geológicos - Madrid 67: 443-453.
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Dam J.A. van, Hoek Ostende L.W. van den, Reumer J.W.F. 2011. A new short-snouted shrew from the Miocene of Spain. Geobios 44: 299-307.
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Donovan S.K., Hoek Ostende L.W. van den 2011. Rijckholtia, a new name for the genus Rykholtia Snajdr, 1980 (Arthopoda, Trilobita), preoccupied by Rykholtia Fritsch, 1910 (Mollusca, Gastropoda). Journal of Paleontology 85: 804-804.

Prieto J., Hoek Ostende L.W. van den, Böhme M. 2011. Reappearance of Galerix (Erinaceomorpha, Mammalia) at the Middle to Late Miocene transition in South Germany: biostratigraphic and palaeoecologic implications. Contributions to Zoology 80: 179-189.
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2010

Journals SCI, peer-reviewed

Angelone C., Colombero S., Esu D., Giuntelli P., Marcolini F., Pavia M., Trenkwalder S., Hoek Ostende L.W. van den, Zunino M., Pavia G. 2010. Moncucco Torinese, a new post-evaporitic Messinian fossiliferous site from Piedmont (NW Italy). Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 259: 89-104.
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Casanovas-Vilar I., García-Paredes I., Alba D.M., Hoek Ostende L.W. van den , Moyà-Solà S. 2010. The European Far West: Miocene mammal isolation, diversity and turnover in the Iberian Peninsula. Journal of Biogeography 37: 1079-1093.
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Zijlstra J.S., Madern P.A., Hoek Ostende L.W. van den 2010. New genus and two new species of Pleistocene oryzomyines (Cricetidae: Sigmodontinae) from Bonaire, Netherlands Antilles. Journal of Mammalogy 91: 860-873.
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Meijer H.J.M., Hoek Ostende L.W. van den, Bergh G.D. van den, Vos J. de 2010. The fellowship of the hobbit: the fauna surrounding Homo floresiensis. Journal of Biogeography 37: 995-1006.
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Journals non-SCI, peer-reviewed

Hoek Ostende L.W. van den, Donovan S.K. 2010. Cor Winkler Prins, the silent force. Scripta Geologica: Special Issue 7: 1-11.
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Edited books

Donovan S.K., Hoek Ostende L.W. van den (eds.). 2010. Palaeontological papers in honour of Dr. Cor F. Winkler Prins. Scripta Geologica Special Issue Scripta Geologica Special Issue, 7 7: 183 p. ISBN 9789081643610.

Wesselingh F.P., Moerdijk P.W., Hoek Ostende L.W. van den (eds.). 2010. Fossiele schelpen van de Nederlandse Kust. 332 p. NCB Naturalis, Leiden. ISBN 9789050113427.

2009

Journals SCI, peer-reviewed

Rijsdijk K.F., Hume J.P., Bunnik F., Florens F.B.V., Baider C., Shapiro B., Plicht J. van der, Janoo A., Griffiths O., Hoek Ostende L.W. van den, Cremer H., Vernimmen T., Louw P.G.B. de, Bholah A., Saumtally S., Porch N., Haile J., Buckley M., Collins M., Gittenberger E. 2009. Mid-Holocene vertebrate bone Concentration-Lagerstatte on oceanic island Mauritius provides a window into the ecosystem of the dodo (Raphus cucullatus). Quaternary Science Reviews 28: 14-24.
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Hoek Ostende L.W. van den, Furio M., Garcia-Paredes I. 2009. New data on Paenelimnoecus from the middle Miocene of Spain support the shrew subfamily Allosoricinae. Acta Palaeontologica Polonica 54: 159-164.

Bergh G.D. van den, Meijer H.J.M., Awe R.D., Morwood M.J., Szabo K., Hoek Ostende L. W. van den, Sutikna T., Saptomo E.W., Piper P.J., Dobney K.M. 2009. The Liang Bua faunal remains: a 95 k.yr. sequence from Flores, East Indonesia. Journal of Human Evolution 57: 527-537.
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Donovan S.K., Hoek Ostende L.W. van den 2009. What's in a name? Or a namer's name? A reply to Dubois (2008). Biological Journal of the Linnean Society 96: 709-711.
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Bergh G.D. van den, Meijer H.J.M., Due R.A., Morwood M.J., Szabó, K., Hoek Ostende L.W. van den, Sutikna .T, Saptomo E.W., Piper P.J., Dobney K.M. 2009. The Liang Bua faunal remains: a 95k.yr. sequence from Flores, East Indonesia. Journal of Human Evolution 57: 527-37.
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Hoek Ostende L.W. van den, Meijer H.J.M., Geer A.A.E. van der 2009. A bridge too far - Comment on "Processes of island colonization by Oligo-Miocene land mammals in the central Mediterranean: New data from Scontrone (Abruzzo, Central Italy) and Gargano (Apulia, Southern Italy)" by PPA Mazza and M. Rustioni [Palaeogeography, Palaeoclimatology, Palaeoecology 267 (2008) 208-215]. Palaeogeography, Palaeoclimatology, Palaeoecology 279: 128-130.
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Callomon J.H., Donovan S.K., Hoek Ostende L.W. van den 2009. Zenostephanus, a new name for the genus Xenostephanus Arkell and Callomon, 1963 (Mollusca, Cephalopoda), preoccupied by Xenostephanus Simpson, Minoprio and Patterson, 1962 (Mammalia). Palaeontology 52: 671-672.

Hoek Ostende L.W. van den, Meijer H.J.M., Geer A.A.E van der 2009. A bridge too far, reply to “Processes of island colonization by Oligo-Miocene land mammals in the central Mediterranean: New data from Scontrone (Abruzzo, Central Italy) and Gargano (Apulia, Southern Italy)” by Mazza P. and Rustioni M. 2008. Palaeogeography, Palaeoclimatology, Palaeoecology 267
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Journals non-SCI, peer-reviewed

Bendukidze O.G., Bruijn H. de, Hoek Ostende L.W. van den 2009. A revision of Late Oligocene associations of small mammals from the Aral Formation (Kazakhstan) in the National Museum of Georgia, Tbilissi. Palaeodiversity 2: 343-377.

Hoek Ostende L.W. van den, Reumer J.W.F., Doukas C.S. 2009. The nature of the fossil record of Neogene insectivores. Hellenic Journal of Geosciences 44: 117-124.

2008

Journals SCI, peer-reviewed

Zijlstra J.S., Hoek Ostende L.W. van den, Due R.A. 2008. Verhoeven's giant rat of Flores (Papagomys theodorverhoeveni, Muridae) extinct after all?. Contributions to Zoology 77: 25-31.

Mayhew D.F., Dieleman F.E., Boele J., Verhaard L., Hoek Ostende L.W. van den 2008. Mimomys hajnackensis from the Pliocene of the Netherlands. Netherlands Journal of Geosciences - Geologie en Mijnbouw 87: 181-188.

Journals non-SCI, peer-reviewed

Morales J., Pozo M., Silva P.G., Domingo M.S., López-Antoñanzas R., Álvarez Sierra M.A., Antón M., Martín Escorza C., Quiralte V., Salesa M.J., Sánchez I.M., Azanza B., Calvo J.P., Carrasco P., García-Paredes I., Knoll F., Hernández Fernández M., Hoek Ostende L. van den, Merino L., Meulen A.J. van den, Montoya P., Peigné S., Peláez-Campomanes P., Sánchez-Marco A., Turner A., Abella J., Alcalde G.M., Andrés M., DeMiguel D., Cantalapiedra J.L., Fraile S., García Yelo B.A., Gómez Cano A.R., López Guerrero P., Oliver Pérez A., Siliceo G. 2008. El sistema de yacimientos de mamíferos miocenos del Cerro de los Batallones, Cuenca de Madrid: estado actual y perspectivas. Seminario de Paleontología de Zaragoza 8: 41-117.

2007

Journals SCI, peer-reviewed

Bruijn H. de, Hoek Ostende L.W. van den, Donovan S.K. 2007. Mirrabella, a new name for the genus Mirabella De Bruijn et al., 1987 (Mammalia), preoccupied by Mirabella Emeljanov, 1982 (Insecta). Contributions to Zoology 76: 279-280.

Gassó Miracle M.E., Hoek Ostende L.W. van den, Arntzen J.W. 2007. Type specimens of amphibians in the National Museum of Natural History, Leiden, the Netherlands. Zootaxa 1482: 25-68.

García-Paredes I., Ostende L.W. van den Hoek 2007. A nomenclatural note on the Miocene dormouse Quercomys parsani Daams, 1991. Contributions to Zoology 76: 205-206.

Edited books

Hoek Ostende L.W. van den, Wesselingh F. (eds.). 2007. De mineralen van Nederland. 104 p. Naturalis, Leiden. ISBN 9789063910037.

Chapters in books

López Guerrero P., Oliver Pérez A., Álverez Sierra M.A., Garcia Paredes I., Hoek Ostende L.W. van den, Peláez-Campomanes de Labra P. 2007. Paleontológia del yacimiento de vertebrados Aragoniense de Casa Montero (Madrid). 247-254: Imprenta Provincial, Cuenca. ISBN 8493557374.

2006

Journals SCI, peer-reviewed

Dam J.A. van, Abdul Aziz H., Alvarez Sierra M.A., Hilgen F.J., Hoek Ostende L.W. van den, Lourens L.J., Mein P., Meulen A.J. van der, Pelaez-Campomanes P. 2006. Long-period astronomical forcing of mammal turnover. Nature 443: 687-691.
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Smith R., Hoek Ostende L.W. van den 2006. A new heterosoricid shrew from the lowermost Oligocene of Europe. Acta Palaeontologica Polonica 51: 381-384.

Journals non-SCI, peer-reviewed

Alvarez Sierra M.A., Garcia Paredes I., Hoek Ostende L. van den, Meulen A.J. van der, Peláez-Campomanes P., Sevilla P. 2006. The Middle Aragonian (Middle Miocene) micromammals from La Retama (Intermediate Depression, Tagus Basin) Province of Cuenca, Spain. Estudios Geológicos - Madrid 62: 401-428.

Bruijn H. de, Mayda S., Hoek Ostende L.W. van den, Kaya T., Saraç G. 2006. Small mammals from the Early Miocene of Sabuncubeli (Manisa, S.W. Anatolia, Turkey). Beiträge zur Paläontologie 30: 57-87.

Doukas C.S., Hoek Ostende L.W. van den 2006. Insectivores (Erinaceomorpha, Soricomorpha; Mammalia) from Karydia and Komotini (Thrace, Greece; MN 4/5). Beiträge zur Paläontologie 30: 109-131.

Hoek Ostende L.W. van den, Fejfar O. 2006. Erinaceidae and Talpidae (Erinaceomorpha, Soricomorpha, Mammalia) from the Lower Miocene of Merkur-Nord (Czech Republic, MN 3). Beiträge zur Paläontologie 30: 175-203.

Hoek Ostende L.W. van den, Bergh G. van der, Rokus A.D. 2006. First fossil insectivores from Flores. Hellenic Journal of Geosciences 41: 67-72.

2005

Journals SCI, peer-reviewed

Storm P., Aziz F., Vos J. de, Kosasih D., Baskoro S., Ngaliman, Hoek Ostende L.W. van den 2005. Late Pleistocene Homo sapiens in a tropical rainforest fauna in East Java. Journal of Human Evolution 49: 536-45.
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Journals non-SCI, peer-reviewed

Hoek Ostende L.W. van der 2005. The case of the shrinking mammoth. Mitteilungen der Kommission für Quartärforschung der Österreichische Akademie der Wissenschaften 14: 59-65.

Edited books

Hoek Ostende L.W. van den, Doukas C.S., Reumer J.W.F. (eds.). 2005. The Fossil Record of the Eurasian Neogene Insectivores (Erinaceomorpha, Soricomorpha, Mammalia), Part I. Scripta Geologica Special Issue 5 300 p. National Museum of Natural History, ISBN 9789073239968.

2003

Journals non-SCI, peer-reviewed

Hoek Ostende L.W. van den 2003. Insectivores (Lipotyphla, Mammalia) from the Ramblian of the Daroca-Calamocha area. 1: 281-310.

Reumer J.W.F., Hoek Ostende L.W. van den 2003. Petauristidae and Sciuridae (Mammalia, Rodentia) from Tegelen, Zuurland, and the Maasvlakte (the Netherlands). 10: 455-467.

Bruijn H. de, Hoek Ostende L.W. van den, Kristkoiz-Boon E., Rummel M., Theocharopoulos C., Ünay E. 2003. Rodents, lagomorphs and insectivores from the middle Miocene hominoid locality of Çandir (Anatolia). CFS Courier Forschungsinstitut Senckenberg 240: 51-87.

Hoek Ostende L.W. van den 2003. Riddleria atecensis nov. gen. nov. sp., a peculiar erinaceid (Erinaceomorpha, Mammalia) from the Lower Miocene of Spain. Beiträge zur Paläontologie 28: 1-11.

2001

Journals SCI, peer-reviewed

Hoek Ostende L.W. van den 2001. A revised generic classification of the Galericini (Insectivora, Mammalia) with some remarks on their palaeobiogeography and phylogeny. Geobios 34: 681-695.

Hoek Ostende L.W. van den 2001. Insectivore faunas from the Lower Miocene of Anatolia - Part 6: Crocidosoricinae (Soricidae). Scripta Geologica 122: 47-81.

Journals non-SCI, peer-reviewed

Hoek Ostende L.W. van den 2001. Insectivore faunas from the Lower Miocene of Anatolia - Part 5: Talpidae. Scripta Geologica 122: 1-45.

Hoek Ostende L.W. van den 2001. Insectivore faunas from the Lower Miocene of Anatolia - Part 7: The Kargi assemblages. Scripta Geologica 122: 83-99.
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Hoek Ostende L.W. van den 2001. Insectivore faunas from the Lower Miocene of Anatolia - Part 8: Stratigraphy, palaeoecology, palaeobiogeography. Scripta Geologica 122: 101-122.

1997

Journals SCI, peer-reviewed

Hoek Ostende L.W. van den 1997. Insectivore faunas from the Lower Miocene of Anatolia - Part 4: The genus Desmanodon (Talpidae) with the description of a new species from the Lower Miocene of Spain. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen 100: 27-65.
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1995

Journals SCI, peer-reviewed

Hoek Ostende L.W. van den 1995. Insectivore faunas from the Lower Miocene of Anatolia - Part 2: Dinosorex (Heterosoricidae). Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen 98: 1-18.
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Hoek Ostende L.W. van den 1995. Insectivore faunas from the Lower Miocene of Anatolia - Part 3: Dimylidae. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen 98: 19-38.
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1992

Journals SCI, peer-reviewed

Bruijn H. de, Ünay E., Hoek Ostende L.W. van den, Saraç G. 1992. A new association of small mammals from the lowermost Lower Miocene of Central Anatolia. Geobios 25: 651-670.

Hoek Ostende L.W. van den 1992. Insectivore faunas from the Lower Miocene of Anatolia - Part 1: Erinaceidae. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen: Biological, Chemical, Geological 95: 437-467.
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1989

Journals SCI, peer-reviewed

Hoek Ostende L.W. van den, Rümke C.G., Hogeweg P. 1989. The use of time-constrained minimal spanning subtrees in the reconstruction of the phylogeny of the European desmanine moles (Desmaninae, Talpidae, Insectivora). Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen - Series B: Palaeontology, Geology, Physics and Chemistry 92: 47-60.

Journals non-SCI, peer-reviewed

Hoek Ostende L.W. van den 1989. The Talpidae (Insectivora, Mammalia) of Eggingen-Mittelhart (Baden-Württemberg, (F.R.G.) with special reference to the Paratalpa-Desmanodon lineage. Stuttgarter Beiträge zur Naturkunde, Serie B 152: 1-29.

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