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Jeremy Miller

Dr. J. (Jeremy) Miller, Senior researcher - Terrestrial Zoology, Spiders


Email: jeremy.miller@naturalis.nl
Phone: +31 (0)71-75 17 379
Room number: C.03.12, Darwinweg 2
“Unlock the potential of biodiversity data”

I am an advocate for the free and open exchange of scientific data. I take this position not only because (through public financial support) vast quantities of data rightly belong in the public domain, but because scientific power depends on independent verifiability. Evolving technologies, data standards, and norms of scientific practice are encouraging open access publishing practices and the aggregation of primary data formatted according to community standards. These trends have exciting potential benefits for science and society.


I hold positions as senior researcher at the Naturalis Biodiversity Center, research associate at the California Academy of Sciences, and member of Plazi, a Swiss organization developing persistent and openly accessible digital taxonomic literature. I am a subject editor for Zootaxa (amaurobioid spiders) and ZooKeys (spiders), and a senior editor of Biodiversity Data Journal.

Postdoctoral fellowships

Postdoctoral Fellow, California Academy of Sciences, 2007. Advisor: Charles Griswold. Funded by "Biotic Survey of the Gaoligongshan, a Biodiversity Hotspot in Western Yunnan, China," National Science Foundation award BSI 0103795 (P.I.s N. Jablonski and P. Fritsch).

California Academy of Sciences, Schlinger Foundation Postdoctoral Fellowship in Systematic Entomology, California Academy of Sciences, 2005-2007. Advisor: Charles Griswold. Systematics of Widow Spiders (Araneae, Theridiidae, Latrodectus): A Total Evidence Approach.

National Museum of Natural History, Smithsonian Institution, 2004-2005. Advisors: Jonathan Coddington and Dr. Ted Schultz. Molecular Phylogeny and Taxonomy of Widow Spiders (Araneae, Theridiidae, Latrodectus).

National Museum of Natural History, Smithsonian Institution, 2003-2004. Advisor: Jonathan Coddington. Funded by "Assembling the Tree of Life: Phylogeny of Spiders," National Science Foundation award DEB 0228699 (P.I.s W. Wheeler, L. Prendini, J. Coddington, G. Hormiga and P. Sierwald).


The George Washington University (GWU), Department of Biological Sciences, Washington, D.C., 1996-2003, Ph.D. Co-advisors: Gustavo HormigaJonathan CoddingtonDissertation: Progress in the systematics of erigonine spiders (Araneae, Linyphiidae, Erigoninae).

Western Carolina University (WCU), Department of Biology in Cullowhee, North Carolina, 1994-1996, M.S. Advisor: Frederick A. Coyle. Thesis: Systematics of the erigonine spider genus Sisicottus (Araneae, Linyphiidae).

The Evergreen State College (TESC), Olympia, Washington, 1989-1994, B.A., B.S.

The Northwest School, Seattle, Washington, 1985-1989.


Research interest

I  use a combination of morphology and DNA sequence data to investigate patterns of biodiversity in spider communities. My work in spider taxonomy follows an open access cybertaxonomic model. In addition to traditional publications, elements of my taxonomic work are distributed online to an assortment of databases and resources. With primary data aggregated according to community standards, it becomes possible to reuse and recombine data without restrictions. I use an XML markup approach to visualize and aggregate primary data from legacy taxonomic publications, suggesting new possibilities for legacy biodiversity data.


Current research topics

Fast forward to:

Cyberdiversity and the spiders of Vietnam

My collaborators and I established one-hectare plots in three national parks in Vietnam. At each location, we conducted a structured inventory of spiders. Images and DNA barcodes from the inventory have been posted online (www.digitalSpiders.org). In what I call the cyberdiversity approach, data sharing can facilitate reconciliation of inventory results between independent studies where current taxonomic knowledge is incomplete, and contribute to an understanding of how diverse communities change over space and time (Miller et al. 2014). 

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From taxonomic literature to cybertaxonomic content

Biodiversity is one of the most information-rich fields of human knowledge, but advances in basic cybertaxonomic infrastructure have only recently provided the tools to organize biodiversity information in ways that respond to a wide range of user groups, including ecologists, land managers, and interested citizens, not to mention the benefits of readily accessible information to the global taxonomic community. The key to unlocking this potential is semantic tagging of both prospective and retrospective taxonomic literature (Miller et al. 2012). I am working with Plazi to develop fine-grained XML markup of legacy taxonomic literature, which allows primary data to be aggregated, linked, and visualized in novel and useful ways. 

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Cybertaxonomic publishing model

Taxonomic descriptions are based on primary specimen data, including the place and time where specimens were collected, images, and DNA sequences. When structured according to community standards (e.g., TDWG) and made available to online databases, these data can be aggregated, distributed, and built upon by others. Open access publishing improves the availability of taxonomic research products internationally within and beyond the immediate taxonomic community. Contributing primary data to online databases and biodiversity resources (e.g., Morphbank, Encyclopedia of Life, GBIFDryad, ZooBank) facilitates aggregation of data from multiple sources, contributing to an expanding picture of life on Earth. 

My taxonomic contributions cover a range of spider lineages, including Eresidae (Miller et al. 2012: Dryad, KML for Google Earth, Nexus aligned sequence matrix), Penestomidae (Miller et al. 2010: GBIFKML for Google Earth), Ctenidae (Miller and Rahmadi 2012GBIF, ZooBank), and the symphytognathoid families (Miller et al. 2014Miller et al. 2009: GBIFspecimen data in ExcelKML for Google Earth). 

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DNA barcoding

I collaborate with the Naturalis Barcoding Facility on various projects. One of these investigated a museum collection as a source of material for DNA barcoding spiders. Several factors were considered, including time since collection, body size, phylogenetic distance, and extraction method. We found that body size and age of oldest successful DNA barcode are significantly correlated after factoring out phylogenetic effects using independent contrasts analysis. Nondestructive extractions significantly extended the DNA barcoding shelf life of museum specimens, especially small-bodied species. This study will help us to more efficiently use legacy collections for spider DNA barcoding projects and predict when fresh specimens will be required (Miller et al. 2013: Dryad). 

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Biodiversity of the Gaoligongshan

Since 1998, the California Academy of Sciences has lead an international interdisciplinary biotic inventory in the Gaoligongshan (or Gaoligong mountains) of western Yunnan, China. The Gaoligongshan is located in the heart of one of the world's biodiversity "hotspots". "Hotspots" are areas of extreme biodiversity and endemism under severe threat of habitat destruction. But the hotspot designation itself, while an extremely successful concept for the conservation community, has been criticized for over-reliance on data from vascular plants and tetrapod vertebrates. The hotspot designation criteria were originally selected in part because these taxa are relatively well known and well sampled worldwide, but it is not clear that they are representative of the biota as a whole because sampling and knowledge about the most diverse groups on Earth tends to be incomplete. Our inventory emphasized vascular plants and tetrapods (hotspot taxa) as well as fish, non-vascular plants (bryophytes), and two arthropod groups. From these data, we will be able to determine whether hotspot taxa are representative of a broader spectrum of the biota for conservation assessment and provide a model for assessing biodiversity of both well known and poorly known groups simultaneously.

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Phylogeny of spiders

I have used molecular sequence data to investigate questions about the phylogenetic affinities of several groups of entelegyne spiders (Miller et al. 2010). Penestomus was first described from females only and placed in the family Eresidae. Discovery of the male decades later brought surprises, especially in the anatomy of the male palp which features (among other things) a retrolateral tibial apophysis (RTA). The presence of an RTA is synapomorphic for a large clade of spiders exclusive of Eresidae. A molecular data matrix based on four loci was constructed to test two alternative hypotheses: 1) Penestomus are eresids and the RTA is convergent, or 2) Penestomus belong within the RTA clade. Our analysis supported Penestomus as an RTA clade lineage not close to Eresidae. As a result, Penestomus was removed from Eresidae and promoted to family rank (Penestomidae). In addition, we transferred the subfamily Coelotinae from the Amaurobiidae to the Agelenidae. A taxonomic revision of Penestomus (Miller et al. 2010) with cybertaxonomic enhancements (including registration of new species in ZooBankEncyclopedia of Life species pages, and specimen records explorable in Google Earth) complements this phylogenetic work.

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Repeated evolution of male sacrifice behavior

While it is true that cannibalism during mating occurs with more or less frequency in many spider species, the notion that black widow spiders always eat their mates is a myth. However, males of the Australian widow spiderLatrodectus hasselti exhibit remarkable mating behavior: during copulation, males execute a somersault that places their abdomen near the mouth of the female, frequently resulting in the death of the male. This is an example of male sacrifice behavior. Male sacrifice behavior, defined as either complicity in cannibalism or spontaneous death associated with copulation, is not limited to widow spiders. An analysis using phylogeny-based statistics suggests that a series of evolutionary prerequisites are associated with the evolution of male sacrifice behavior in spiders, including extreme sexual size dimorphism, accumulation of multiple males in the female web (an indicator of a male-biased effective sex ratio), and mutilation of the male genitalia during mating (Miller 2007).

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Donat Agosti, Plazi

Maydianne Andrade, University of Toronto, Scarborough

Kevin K. Beentjes, DNA barcoding facilityNaturalis Biiodiversity Center

Terry Catapano, Columbia University, New York

Pingping ChenNaturalis Biiodiversity Center

Torsten Dikow, Smithsonian National Museum of Natural History

Charles Griswold, Department of Entomology, California Academy of Sciences

Peter van Helsdingen, Naturalis Biiodiversity Center and European Invertebrate Surevy

Dave KavanaughDepartment of EntomologyCalifornia Academy of Sciences

Sarah MaunsellStability of Altered Forest Ecosystem (SAFE) Project

Josh Miller, University of Cincinnati

Lyubomir Penev, Pensoft Publishing

Pham Dinh Sac, Vietnam Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology

Martin Ramírez, Museoi Argentino de Ciencias Naturalis Bernardino Rivadavia

Guido Sauter, Plazi

Menno Schilthuizen, Naturalis Biiodiversity Center

Soraya SierraNaturalis Biiodiversity Center

Rutger VosNaturalis Biiodiversity Center


Loans and collection visits can be arranged through our collection department



I participate in courses offered through Leiden University including the Master's level orientation "Evolution, Biodiversity and Conservation", the Tropical Ecology Field course, and the undergraduate Biodiversity course. Stage opportunities are available for students interested in quantitative biodiversity, community ecology, and cybertaxonomy. 

Supervision BSc Students

2014 Adrià Bellvert Bantí, University of Girona, External Academic Training
2010 Manon de Waard, University of Leiden, internship
Rense Schelfhorst, University of Leiden, internship

Supervision MSc Students

2012 Phung Thi Hong Luong, Institute of Ecology and Biological Resources, Vietnamese Academy of Science and Technology, Martin Fellow
2012 Youri Lammers, University of Leiden, internship
2010 Mohammad Marhabaie, University of Isfahan, Iran, Martin Fellow
Available student projects

Public outreach

Selected Popular Features and Presentations

Cannibalized and Loving It. Naturalis After Dark: Nature's Peepshow. 5 June 2014. Link

Mooiste spinnen van Europa komen bovengronds in online bibliotheek. 12 June 2012. Link

Rahman, R. Meet the new official Lou Reed spider breed: Just as cuddly as the man himself! EW.com. 24 May 2012. Link

Prostak, S. New Genus of Velvet Spider Named after Lou Reed. Sci-news.com. 23 May 2012. Link

Velvet spiders emerge from underground in new cybertaxonomic monograph. Pensoft news. 23 May 2012. Link

Accessible Biodiversity Knowledge and the World Wide Web. Naturalis After Dark. 28 February 2012. Explore the Prezi

Interview. In: Huang, C. Redback spider fears stalk the Emirates. The National. 18 December 2010. Link

Interview. In: Dolgin, E. 2009. Lazy male spiders avoid dinner date. Nature. 21 October 2009. doi:10.1038/news.2009.1020. Link

Interview. Black Widows - Part 2. Science in Action, California Academy of Sciences podcast. 3 September 2007.

Interview. Black Widows - Part 1. Science in Action, California Academy of Sciences podcast. 27 August 2007.

Interview. Silk Triplines. Science in Action, California Academy of Sciences podcast. 20 August 2007.

Interview. In: Suicide mating. Pp. 16-17. California Academy of Sciences: Member Publication, Summer 2007.

Interview, photography. In: Perlman, D. Academy of Sciences. Spitting spider's surprise. Unlike its brethren, this new discovery has a nurturing side. San Francisco Chronicle. 22 July 2006. Link

Interview, photography. In: Spiders that hunt together stay together. Pp. 19-21. California Academy of Sciences: Member Publication, Fall 2006.

Interview, cover photo and additional photography. In: Tiny Assassins. Pp. 3-4. California Academy of Sciences: Member Publication, Summer 2006.

Tiny Assassins. California Academy of Sciences, Science Now. 2006. Link

California Academy of Sciences Annual Halloween Party and Benefit, San Francisco, California, 2006

Tagliaferro, L. 2004. Spiders and Their Webs. Animal Homes series. J. Miller, consultant. Capstone Press, Mankato, Minnesota.

Insect Expo, National Museum of Natural History, Washington D.C., 2003.

Bugfest, National Mall, Washington D.C., 1997, 2000.

"Spellbinding Spiders," a public presentation at the Mountain Heritage Center, Cullowhee, North Carolina, 1995.

"Spiders!" an arachnological experience for preschool-age naturalists at the Jackson County Public Library, Sylva, North Carolina, 1995.



Journals SCI, peer-reviewed

Schilthuizen M., Vairappan C.S., Slade E.M., Mann D.J., Miller J.A. 2015. Specimens as primary data: museums and ‘open science’. 30: 237-238.
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Journals non-SCI, peer-reviewed

Miller J.A., Georgiev T., Stoev P., Sautter G., Penev L. 2015. Corrected data re-harvested: curating literature in the era of networked biodiversity informatics. Biodiversity Data Journal 3: e4552.
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Miller J.M., Agosti D., Penev L., Sautter G., Georgiev T., Catapano T., Patterson D., King D., Pereira S., Vos R.A., Sierra S. 2015. Integrating and visualizing primary data from prospective and legacy taxonomic literature. Biodiversity Data Journal 3: 1-33.
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Claessens J., Beentjes K.K., Heijerman T., Miller J., Gravendeel, B. 2015. Beobachtungen von Miarus campanulae als Bestäuber von Cephalanthera rubra.: 77-87.


Journals SCI, peer-reviewed

Yang C., Wang X., Miller J.A., Blécourt M. de, Ji Y., Yang C., Harrison R.D., Yu D.W. 2014. Using metabarcoding to ask if easily collected soil and leaf-litter samples can be used as a general biodiversity indicator. Ecological Indicators 46: 379-389.
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Kuntner M., Pristovšek U., Cheng R-C., Li D., Zhang S., Tso I-M., Liao C-P., Miller J.A., Kralj-Fišer S. 2014. Eunuch supremacy: evolution of post-mating spider emasculation. Behavioral Ecology and Sociobiology 69: 117-126.
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Miller J.A., MillerJ.H., Pham D-S., Beentjes K.K. 2014. Cyberdiversity: improving the informatic value of diverse tropical arthropod inventories. PLOS ONE 9: e115750.
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Ramírez M.J., Grismado C.J., Labarque F.M., Izquierdo M.A., Ledford J.M., Miller J.A., Haddad C.R., Griswold C.E. 2014. The morphology and relationships of the walking mud spiders of the genus Cryptothele (Araneae: Zodariidae). Zoologischer Anzeiger 253: 382-393.
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Journals non-SCI, peer-reviewed

Miller J.A., Schilthuizen M., Lilliendahl Burmester J., Graaf L. van der, Merckx V., Jocqué M., Kessler P.J.A., Fayle T.M., Breeschoten T., Broeren R., Bouman R., Chua W.-J., Feijen F., Fermont T., Groen K., Groen M., Kil N.J.C., Laat H.A. de, Moerland M.S., Moncoquet C., Panjang E., Philip A.J., Roca-Eriksen R., Rooduijn B., Santen M. van, Swakman V., Evans M.N., Evans L.J., Love K., Joscelyne S.H., Tober A.V., Wilson H.F., Ambu L.N., Goossens B. 2014. Dispatch from the field: ecology of ground-web-building spiders with description of a new species (Araneae, Symphytognathidae). Biodiversity Data Journal 2: 1-13.
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Vos R.A., Biserkov J.V., Balech B., Beard N., Blissett M., Brenninkmeijer C., Dooren T. van, Eades D., Gosline G., Groom Q.J., Hamann T.D., Hettling H., Hoehndorf R., Holleman A., Hovenkamp P., Kelbert P., King D., Kirkup D., Lammers Y., DeMeulemeester T., Mietchen D., Miller J.A., Mounce R., Nicolson N., Page R., Pawlik A., Pereira S., Penev L., Richards K., Sautter G., Shorthouse D.P., Tähtinen M., Weiland C., Williams A.R., Sierra S. 2014. Enriched biodiversity data as a resource and service. Biodiversity Data Journal 2: 1-15.
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Journals SCI, peer-reviewed

Miller J.A., Beentjes K.K., Helsdingen P. van, IJland S. 2013. Which specimens from a museum collection will yield DNA barcodes? A time series study of spiders in alcohol. 365: 245-261.
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Journals non-SCI, peer-reviewed

Miller J., Belgers J., Beentjes K., Zwakhals K., Helsdingen P. van 2013. Spider hosts (Arachnida, Araneae) and wasp parasitoids (Insecta, Hymenoptera, Ichneumonidae, Ephialtini) matched using DNA barcodes. Biodiversity Data Journal 1: 1-20.
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Smith V., Georgiev T., Stoev P., Biserkov J., Miller J., Livermore L., Baker E., Mietchen D., Couvreur T., Mueller G., Dikow T., Helgen K., Frank J., Agosti D., Roberts D., Penev L. 2013. Beyond dead trees: integrating the scientific process in the Biodiversity Data Journal. Biodiversity Data Journal 1: e995.
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Journals SCI, peer-reviewed

Miller J., Rahmadi C. 2012. A troglomorphic spider from Java (Araneae, Ctenidae, Amauropelma). ZOOKEYS 163: 1-11.
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Miller J., Dikow T., Agosti D., Sautter G., Catapano T., Penev L., Zhang Z.-Q., Pentcheff D., Pyle R., Blum S., Parr C., Freeland C., Garnett T., Ford L.S., Muller B., Smith L., Strader G., Georgiev T., Benichou L. 2012. From taxonomic literature to cybertaxonomic content. BMC BIOLOGY 10: 87.
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Miller J.A., Griswold C.E., Scharff N., Rezac M., Szuts T., Marhabaie M. 2012. The velvet spiders: an atlas of the Eresidae (Arachnida, Araneae). ZOOKEYS 195: 1-144.
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Journals non-SCI, peer-reviewed

IJland S., Helsdingen P.J. van, Miller J. 2012. On some spiders from Gargano, Apulia, Italy. Nieuwsbrief Spined 32: 2-20.
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Journals non-SCI, peer-reviewed

Miller J., Pham D.S. 2011. Landscape biodiversity of tropical forest spider communities in Vietnam (Arachnida: Araneae). Treubia 38: 53-70.

Dumbacher J.P., Miller J. , Flannery M.E., Yang X. 2011. Avifauna of the Gaoligong Shan Mountains of Western China: A hotspot of avian species diversity. Ornithological Monographs 70: 30-63.
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IJland S., Miller J. , Sluijs Y.J. van, Helsdingen P.J. van 2011. Enkele spinnen uit Zuid-Limburg, met Monocephalus castaneipes (Araneae, Linyphiidae) nieuw voor Nederland. Nieuwsbrief Spined 31: 19-22.
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Journals SCI, peer-reviewed

Penev L., Agosti D., Georgiev T., Catapano T., Miller J., Blagoderov V., Roberts D., Smith V.S., Brake I., Ryrcroft S., Scott B., Johnson N.F., Morris R.A., Sautter G., Chavan V., Robertson T., Remsen D., Stoev P., Parr C., Knapp S., Kress W.J., Thompson F.C., Erwin T. 2010. Semantic tagging of and semantic enhancements to systematics papers: ZooKeys working examples. ZOOKEYS 50: 1-16.
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Wang X.-P., Griswold C.E., Miller J.A. 2010. Revision of the genus Draconarius Ovtchinnikov 1999 (Agelenidae: Coelotinae) in Yunnan, China, with an analysis of the Coelotinae diversity in the Gaoligongshan Mountains. ZOOTAXA 2593: 1-127.
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Miller J.A., Griswold C.E., Haddad C.R. 2010. Taxonomic revision of the spider family Penestomidae (Araneae, Entelegynae). ZOOTAXA 2534: 1-36.
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Miller J.A., Carmichael A., Ramirez M.J., Spagna J.C., Haddad C.R., Rezac M., Johannesen J., Kral J., Wang X.-P., Griswold C.E. 2010. Phylogeny of entelegyne spiders: Affinities of the family Penestomidae (NEW RANK), generic phylogeny of Eresidae, and asymmetric rates of change in spinning organ evolution (Araneae, Araneoidea, Entelegynae). MOLECULAR PHYLOGENETICS AND EVOLUTION 55: 786-804.
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Journals SCI, peer-reviewed

Coddington J.A., Agnarsson I., Miller J.A., Kuntner M., Hormiga G. 2009. Undersampling bias: the null hypothesis for singleton species in tropical arthropod surveys. JOURNAL OF ANIMAL ECOLOGY 78: 573-584.
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Journals non-SCI, peer-reviewed

Miller J.A., Griswold C.E., Yin C.M. 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers. ZooKeys 11: 9-195.
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Penev L., Erwin T., Miller J., Chavan V., Moritz T., Griswold C. 2009. Publication and dissemination of datasets in taxonomy: ZooKeys working example. ZooKeys 11: 1-8.
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Helsdingen P.J. van, Miller J., IJland S. 2009. Snelle inventarisatie van het Noordoostpolder (Kuinderbos) in Flevoland (Araneae). Nieuwsbrief Spined 27: 16-17.
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Journals SCI, peer-reviewed

Agnarsson I., Miller J.A. 2008. Is ACCTRAN better than DELTRAN?. CLADISTICS 24: 1032-1038.
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Penev L., Erwin T., Thompson F.C., Sues H.-D., Engel M.S., Agosti D., Pyle R., Ivie M., Assmann T., Henry T., Miller J., Ananjeva N.B., Casale A., Lourenço W., Golovatch S., Fagerholm H.-P., Taiti S., Alonso-Zarazaga M., Nieukerken E. van 2008. ZooKeys, unlocking earth’s incredible biodiversity and building a sustainable bridge into the public domain: From “print-based” to “web-based” taxonomy, systematic, and natural history. ZooKeys 1: 1-17.
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Journals SCI, peer-reviewed

Miller J.A. 2007. Repeated evolution of male sacrifice behavior in spiders correlated with genital mutilation. EVOLUTION 61: 1301-1315.
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Ramirez M.J., Coddington J.A., Maddison W.P., Midford P.E., Prendini L., Miller J., Griswold C.E., Hormiga G., Sierwald P., Scharff N., Benjamin S.P., Wheeler W.C. 2007. Linking of digital images to phylogenetic data matrices using a morphological ontology. SYSTEMATIC BIOLOGY 56: 283-294.
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Miller J.A. 2007. Review of erigonine spider genera in the neotropics (Araneae : Linyphiidae, Erigoninae). ZOOLOGICAL JOURNAL OF THE LINNEAN SOCIETY 149: 1-263.
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Journals non-SCI, peer-reviewed

Miller J.A. 2007. Synaphridae of Madagascar (Araneae: Araneoidea): A new family record for the Afrotropical region. Proceedings of the California Academy of Sciences 58: 21-48.
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Journals SCI, peer-reviewed

Simmons M.P., Zhang L.B., Webb C.T., Reeves A., Miller J.A. 2006. The relative performance of Bayesian and parsimony approaches when sampling characters evolving under homogeneous and heterogeneous sets of parameters. CLADISTICS 22: 171-185.

Journals non-SCI, peer-reviewed

Miller J. 2006. Web-sharing sociality and cooperative prey capture in a Malagasy spitting spider (Araneae: Scytodidae). Proceedings of the California Academy of Sciences 57: 739-750.
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Journals SCI, peer-reviewed

Miller J.A. 2005. Cave adaptation in the spider genus Anthrobia (Araneae, Linyphiidae, Erigoninae). ZOOLOGICA SCRIPTA 34: 565-592.
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Miller J., Agnarsson I. 2005. A redescription of Chrysso nigriceps (Araneae, Theridiidae) with evidence for maternal care. JOURNAL OF ARACHNOLOGY 33: 711-714.
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Miller J.A. 2005. A redescription of Porrhomma cavernicola Keyserling (Araneae, Linyphiidae) with notes on Appalachian troglobites. JOURNAL OF ARACHNOLOGY 33: 426-438.
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Journals SCI, peer-reviewed

Miller J.A. 2004. The genus Brattia beyond South America (Araneae, Linyphiidae). JOURNAL OF ARACHNOLOGY 32: 526-538.
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Miller J.A., Hormiga G. 2004. Clade stability and the addition of data: A case study from erigonine spiders (Araneae: Linyphiidae, Erigoninae). Cladistics 20: 385-442.
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Journals SCI, peer-reviewed

Miller J.A. 2003. Assessing progress in systematics with continuous jackknife function analysis. SYSTEMATIC BIOLOGY 52: 55-65.
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Journals SCI, peer-reviewed

Toti D.S., Coyle F.A., Miller J.A. 2000. A structured inventory of appalachian grass bald and heath bald spider assemblages and a test of species richness estimator performance. JOURNAL OF ARACHNOLOGY 28: 329-345.
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Journals SCI, peer-reviewed

Miller J.A. 1999. Revision and cladistic analysis of the erigonine spider genus Sisicottus (Araneae, Linyphiidae, Erigoninae). JOURNAL OF ARACHNOLOGY 27: 553-603.
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Zujko-Miller J. 1999. On the phylogenetic relationships of Sisicottus hibernus (Araneae, Linyphiidae, Erigoninae). Journal of Arachnology 27: 44-52.
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Journals SCI, peer-reviewed

Miller J.A., Coyle F.A. 1996. Cladistic analysis of the Atypoides plus Antrodiaetus lineage of mygalomorph spiders (Araneae, Antrodiaetidae). JOURNAL OF ARACHNOLOGY 24: 201-213.
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Editorial duties

External links

Online resources and software

Encyclopedia of Life (http://www.eol.org/), 49 species pages contributed.

Morphbank (http://www.morphbank.net/), 2442 images contributed.

ZooBank (http://www.zoobank.org/), 51 nomenclatural acts registered.

Barcode of Life (http://www.boldsystems.org/views/login.php), 695 public sequences contributed. 

Miller, J.A. & P.D. Sac. 2014. digitalSpiders. Beta version 0.6. www.digitalSpiders.org.

Miller, J.A. 2004. Key to the Genera of Erigonine Spiders in the Neotropics (Araneae, Linyphiidae, Erigoninae). Available online at http://www.gwu.edu/~spiders/interactiveKeys.htm. National Museum of Natural History, Smithsonian Institution.

Hormiga, G, J.A. Miller & F. Alverez-Padilla. 2003. LinyGen: Linyphioid Genera of the World (Pimoidae and Linyphiidae), An Illustrated Catalog. Preview Version. Available online athttp://www.gwu.edu/~linygen/.

Zujko-Miller, C. & J.A. Miller. 2002. PEST: Precision Estimated by Sampling Traits. Available online at http://www.gwu.edu/~spiders/pestDocs.htm.

Miller, J.A. & M. Tschapka. 2001. The Bat Flies of La Selva (Diptera: Nycteribiidae, Streblidae). Available online at: http://www.biologie.uni-ulm.de/bio3/Batfly/index.html.

Personal identifiers and profiles

Entomology collection

Visit the Naturalis collections

Loan policy