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Han van Konijnenburg-van Cittert

Prof. Dr. J.H.A. (Han) van Konijnenburg-van Cittert, Research associate - Geology


Email: han.vankonijnenburg@naturalis.nl
Phone: +31 (0)71-7517317
Room number: C01.09, Darwinweg 2

My work on Permian and Triassic floras started in 1991 with the description of a few plant fossils from a Late Triassic flora in Iran. Later I took part in the large project, started by the late Prof. Dr. H.-J. Schweitzer (Bonn, Germany) to describe the whole of the Triassic and Jurassic plant fossils from Iran and Afghanistan. This resulted in a co-authorship of three of the monographs on this flora (Schweitzer et al., 1997, 2000 and 2009; see separate list of publications on Permian and Triassic floras).

I also became involved in the research carried out at the Laboratory of Palaeobotany and Palynology (Utrecht, the Netherlands) on the large extinction event at the end of the Permian and following recovery during the first stage of the Triassic (see separate list of publications).

From 1999 onwards I work on Middle and Late Triassic floras in Northern Italy, dealing in a large number of publications with resp. the Anisian and the Ladinian flora from the Dolomites (with a number of co-authors including Dr. E. Kustatscher whose Msc and PhD theses on these subjects I supervised for a large part) and the Carnian flora of the Bergamasc Alps (a PhD project of Dr. L. Passoni).

Finally, I participated in research papers on Triassic floras in former Yugoslavia, the Netherlands, Germany and Austria (see literature list). I also used data on Permian and Triassic plants in my more general papers dealing with evolution of plant groups etc.



1961-1967 Study Biology University Utrecht.
1967-1970 PhD research; PhD thesis defence: "In situ gymnosperm pollen from the Middle Jurassic of Yorkshire" 2 december 1970 (promotor Prof.F.P.Jonker, Utrecht; co-promotor Prof.T.M. Harris, Reading, Engeland).
1970-present Honorary Scientific researcher University of Utrecht, Laboratory of Palaeobotany and Palynology.
2003-present Extraordinary Professor of Prequaternary Palaeobotany University of Leiden and as such also staff member of the National Herbarium of the Netherlands (Leiden).



In 1999 I received together with K.-P.Kelber (University of Würzburg, Germany) the Remy prize from the Paleobotanical Section of the Botanical Society of America for the paper: ‘Equisetites arenaceus from the Upper Triassic of Germany with evidence for reproductive strategies’.

In 2006, the paper “Abbink, O.A., van Konijnenburg-van Cittert, van der Zwan, C.J. & Visscher, H. 2004. A Sporomorph Ecogroup Model for the Northwest European Upper Jurassic – Lower Cretaceous II: Application to an exploration well from the Dutch North Sea. Neth. J. of Geology/Geologie en Mijnbouw 83(2): 81-92” won the Netherlands Journal for Geology publication price.

Fellowships, Board memberships

  • Fellow of the Linnean Society of London (since 1985)
  • Board of Review of Palaeobotany and Palynology (Elsevier) (since 1989)
  • Board of Acta Palaeobotanica (Krakow, Polen) (since 2000)
  • Board of Scripta geologica (Leiden) (since 2003)
  • Board of the Neues Jahrbuch f. Geologie und Palaeontologie (Stuttgart, Germany) (since 2005)
  • Vice-president of the Treub Maatschappij for research in tropical countries (since 2006)


Research interest

Scientific research projects vary from Early Permian to Late Cretaceous floras, plus the Eocene Azolla bloom in the Arctic Ocean.


palaeobotany, fossil pteridophyta, fossil gymnosperms

Current research topics

  • Inventarisation and determination of the Jongmans collection of plant macrofossils from all ages and from all over the world, but with emphasis on the Carboniferous flora from the Netherlands.
  • Revision of the Early Permian flora from Djambi (Indonesia), together with Dr. Isabel van Waveren and Drs. Menno Booi.
  • Late Cretaceous conifers from the Maastrichtian of the type-area, together with Dr. Van der Ham, National Herbarium, Leiden. 
  • Santonian plant fossils from Limburg, The Netherlands, together with Drs. H. Bosma.

Brachyphyllum patens from the Maastrichtian type area. 

Next to Naturalis, I also work at the Laboratory of Palaeobotany and Palynology, University of Utrecht. There my research interests are:

  • Macropalaeobotany and in situ pollen and spores from the Middle Jurassic of Yorkshire, England.
  • Macropalaeobotany, ecology and in situ pollen and spores from the Late Jurassic flora of Sutherland, Scotland (with Dr. van der Burgh, Utrecht).
  • Late Triassic flora from Bavaria, Germany (with Dr. Kelber, Würzburg).
  • Rhaeto-Liassic flora from Bavaria, Germany (with Dr. Kirchner, München en mr. Schmeißner, Kulmbach).
  • LM, SEM and TEM work on fossil in situ spores and pollen (ferns and gymnosperms); comparison with living material.
  • Ladinian and Anisian floras from the Dolomites (with Dr. Kustatscher, Bolzano and Mr. Wachtler, Innichen, Italy).
  • Permian/Triassic boundary at Bletterbach, Dolomites (with Prof. Kerp, Munster; Dr. Fischer, Munich; Dr. Kustatscher, Bolzano).
  • Azolla remains in the Eocene of the Arctic Ocean (with Prof. Lotter, Prof. Brinkhuis, Dr. van der Burgh, Drs. Barke, all Utrecht and Prof. Collinson, London).

Reconstruction of Neuropteridium elegans from the Anisian of the Dolomites.


In my work at the national Natural History Museum “Naturalis” (Leiden, the Netherlands), I study the Early Permian flora from Jambi (Indonesia) together with Dr. I.M.van Waveren and drs. M. Booi whose PhD on this flora I supervise. Moreover, I supervise the PhD thesis of drs. Hylke Bosma on Late Cretaceous conifers from the Northern Hemisphere, and I work together with dr. Raymond van der Ham (Herbarium) on the Maastrichtian flora of Limburg and surroundings.





Available student projects


Naturalis Repository

In prep

Collinson, M.E. & J.H.A.van Konijnenburg-van Cittert. Scanning and transmission electron microscopy of Azolla primaeva. Review Palaeobotany Palynology.

Looy, C.V., Duijnstee, I.A.P., Grauvogel-Stamm, L. & J.H.A.van Konijnenburg-van Cittert. A new reconstruction of the Triassic lycopsid Pleuromeia sternbergii (Münster) Corda ex Giebel.


Kathleen Bauer, Evelyn Kustatscher, Rainer Butzmann, Thilo C. Fischer, Johanna H.A. van Konijnenburg-van Cittert and Michael Krings. Ginkgophytes from the Upper Permian of the Bletterbach gorge (northern Italy). Rivista Italiana di Paleontologia e Stratigrafia.

In press

Kathleen Bauer, Evelyn Kustatscher, Günther Dütsch, Stefan Schmeißner, Michael Krings & Johanna H. A. van Konijnenburg-van Cittert. Lepacyclotes kirchneri n. sp. (Isoetales, Isoetaceae) aus dem unteren Jura von Oberfranken, Deutschland. Berichte der Naturwissenschaftlichen Gesellschaft Bayreuth.

Recent publications 

Booi, M., van Waveren, I.M. & J.H.A. van Konijnenburg-van Cittert, 2014. Wood anatomical variability in Early Permian ‘Araucarioids’. IAWA Journal 35 (3): 307-331.

Page, C., Collinson, M.E. & J.H.A. van Konijnenburg-van Cittert, 2014. Lygodium hians (Pteridophyta: Schizaeaceae) – a ground-clothing template for the evolution of a modern climbing fern genus. Adansonia Serie 3, vol 36. (1): 1-24.

Kustatscher, E., Bauer, K., Butzmann, R., Fischer, T.C., Meller, B., van Konijnenburg-van Cittert, J.H.A. & H. Kerp, 2014. Sphenophytes, seed ferns and taeniopterids from the Upper Permian of Bletterbach (Dolomites, N-Italy). Review Palaeobotany Palynology 208: 65-82.

Bosma, H.F. & J.H.A. van Konijnenburg-van Cittert, 2014. Revision of the Late Cretaceous conifer genus Moriconia. Palaeobiodiversity and Palaeoenvironments. DOI: 10.1007/s12549-014-0158-8.

J.H.A. van Konijnenburg - van Cittert, 2014. Book review of “Fire on earth: an introduction by Andrew C. Scott, David M.S.J. Bowman, William J. Bond, Stephen J. Pyne and Martin E. Alexander.” Geological Journal Published online in Wiley Online Library (wileyonlinelibrary.com) DOI: 10.1002/gj.2566.

van Konijnenburg-van Cittert, J.H.A., Kustatscher, E., Bauer, K., Pott, C., Schmeissner, S., Dütsch, G. & M. Krings, 2014. A Selaginellites from the Rhaetian of Wüstenwelsberg (Upper Franconia, Germany). Neues Jahrbuch für Geologie und Palaeontologie,  Abhandlungen 272(2): 115-127.

Zijlstra, G. & J.H.A. van Konijnenburg-van Cittert, 2014. A nomenclatural Note on Monanthesia and Bennettites. Review Palaeobotany Palynology 203:1-2.

Welzen, P.C., Strijk, J.S., van Konijnenburg-van Cittert, J.H.A., Nucete, M., Merckx, V.S., 2014. Dated phylogenies of the sister genera Macaranga and Mallotus (Euphorbiaceae): Congruence in Historical Biogeographic patterns? PlosOne. Jan. 17th. DOI:10.1371/journal.pone.0085713.

Ronny Rößler, Marc Philippe, Johanna H.A. Van Konijnenburg-Van Cittert, Stephen McLoughlin, Jakub Sakala, Gea Zijlstra, 2014. Which name should be kept for fossil wood of the Araucarioxylon-type? – Results of a poll. Taxon 63(1): 177-184.

Kustatscher, E. and J.H.A. van Konijnenburg-van Cittert, 2013. Seed ferns from the European Triassic – an overview. New Mexico Museum of Natural History and Science, Bulletin 61: 331-344.

Burgh, J. van der, Collinson, M.E., van Konijnenburg-van Cittert, J.H.A., Barke, J. and H. Brinkhuis, 2013. The freshwater fern Azolla (Azollaceae) from Arctic and Nordic sediments: new species and their stratigraphic distribution. Review Palaeobotany Palynology 194: 50-68. doi:10.1016/j.revpalbo.2013.02.002.

Zijlstra, G., Kustatscher, E. & J.H.A. van Konijnenburg-van Cittert, 2013.Proposal to conserve the name Sphenozamites (Brongn.) Miq. with a conserved type (fossil Cycadophyta: Bennettitales). Taxon 62 (1): 185-186.

Collinson, M.E., Smith, S.Y., van Konijnenburg-van Cittert, J.H.A., Batten, D.J., van der Burgh, J., Barke, J. and Marone, F., 2013. New observations and synthesis of Paleogene heterosporous water ferns. International Journal of Plant Sciences 174 (3): 350-363.

Reichgelt, T., Parker, W.G., van Konijnenburg-van Cittert, J.H.A., Martz, J.W., Kürschner, W.M., 2013. The palynology of the Sonsela Member (Late Triassic, Norian) at Petrified Forest NP, Arizona, USA. Review Palaeobotany Palynology 189: 18-28.

Kustatscher, E., dell’Antonio E., & J.H.A. van Konijnenburg-van Cittert, 2012. The ferns of the late Ladinian (Middle Triassic) flora from Monte Agnello (Dolomites, N-Italy). Acta Palaeontologica Polonica., doi: 10.4202/app.2012.0076.

Kustatscher, E., Heunisch, C. & J.H.A. van Konijnenburg-van Cittert, 2012 Taphonomical implications of the Ladinian macro- and palynofloras of Thale (Germany). Palaios 27(11): 753-764.

Cleal, C.J., Thomas, B.A., van Konijnenburg-van Cittert, J.H.A. & Zijlstra G., 2012. Proposal to change the authorship of Calamites, nom. cons. (fossil Sphenopsida) and to delete Calamitis, nom. rej.. Taxon 61 (4): 884-885.

Nucete, M., van Konijnenburg-van Cittert, J.H.A. and P.C. van Welzen, 2012. Fossils and Palaeontological distributions of Macaranga and Mallotus (Euphorbiaceae). Palaeogeography, Palaeoclimatalogy, Palaeoecology 353-355: 104-115.

Kustatscher, E., Kelber, K.-P. & J.H.A. van Konijnenburg-van Cittert, 2012. Danaeopsis Heer ex Schimper 1869 and its European Triassic species. Review Palaeobotany and Palynology 183: 32-49. 

Bosma, H., Kunzmann, L., Kvacek, J. & J.H.A.van Konijnenburg-van Cittert, 2012. Revision of the Cretaceous conifer genus Cunninghamites. Review Palaeobotany Palynology 182: 20-31.

Kustatscher, E., van Konijnenburg-van Cittert, J.H.A., Bauer, K., Butzmann, R., Meller, B., Fischer, T.C., 2012. A new macroflora from the Upper Permian of Bletterbach (Dolomites, N. Italy). Review Palaeobotany Palynology 182: 1-13.

Judith Barke, Johan van der Burgh, Johanna H. A. van Konijnenburg-van Cittert, Margaret E. Collinson, Martin A. Pearce, Jonathan Bujak, Claus Heilmann-Clausen, Eveline N. Speelman, Monique M.L. van Kempen, Gert-Jan Reichart, André F. Lotter, and Henk Brinkhuis, 2012. Coeval Eocene blooms of the fresh water fern Azolla in and around the Arctic and Nordic Seas. Palaeogeography, Palaeoclimatalogy, Palaeoecology 337-338: 108-119.

Zavialova, N. E. & J.H.A.van Konijnenburg-van Cittert, 2012. Exine ultrastructure of pollen of Androstrobus prisma Thomas et Harris 1960 from the Jurassic of Yorkshire. Review of Palaeobotany and Palynology 173: 15-22.

J.H.A. van Konijnenburg - van Cittert, 2012. Book review of “The Terrestrialization process: Modelling complex interactions at the Biosphere-Geosphere interface.“ Geological Journal 47: 356. on line: DOI: 10.1002/gj.1297.

Kustatscher, E., Pott, C. & J.H.A. van Konijnenburg-van Cittert, 2011. Scytophyllum waehneri (Stur) nov. comb., the correct name for Scytophyllum persicum (Schenk)Kilpper, 1975. Zitteliana A 51: 9-18.

Van der Ham, R. W. J. M., van Konijnenburg-van Cittert, J.H.A., Kieft, B. N. & A. Walsmit Sachs, 2011. Mosacaulis spinifer gen. et sp. nov.: a Late Cretaceous marine lycopsid. Review of Palaeobotany and Palynology 168: 51-67.

Zavialova, N. E. & J.H.A.van Konijnenburg-van Cittert, 2011. Exine ultrastructure of in situ peltasperm pollen from the Rhaetian of Germany and its implications. Review of Palaeobotany and Palynology 168: 7-20.

Kustatscher, E. & J.H.A. van Konijnenburg-van Cittert, 2011. The ferns of the Middle Triassic flora from Thale. Neues Jahrbuch für Geologie und Palaeontologie,  Abhandlungen 261(2): 209-248.

Kustatscher, E., Pott, C. & J.H.A. van Konijnenburg-van Cittert, 2011. A contribution to the knowledge of the Triassic fern genus Symopteris. Review of Palaeobotany and Palynology 165: 41-60.

Zavialova, N.E., Kustatscher, E. & Johanna H.A. van Konijnenburg-van Cittert, 2010. Spore ultrastructure of Selaginellites leonardii. Geo.Alp. 7: 1-18.

Gea Zijlstra, Evelyn Kustatcher & Han van Konijnenburg-van Cittert, 2010. Proposal to conserve the name Danaeopsis Heer ex Schimp. 1869 (fossil Pteridophyta) against Marantacea Jaeger 1827 (fossil Pteridophyta) and Danaeopsis C. Presl (recent Pteridophyta). Taxon 59(6): 1987-1988.

J.H.A. van Konijnenburg - van Cittert & Gea Zijlstra, 2010. Nomenclature, typification and correct description of fossil plants. IOP Newsletter 93: 5-9; online: www.palaeobotany.org

J.H.A. van Konijnenburg - van Cittert, 2010. The Early Jurassic male ginkgoalean inflorescence Stachyopitys preslii Schenk and its in situ pollen. Scripta Geologica Special Issue 7: 141-149.

Kustatscher, E. & J.H.A. van Konijnenburg-van Cittert. 2010. Seed ferns and cycadophytes from the Triassic flora of Thale (Germany). Neues Jahrbuch für Geologie und Paläontologie, 258(2): 195-217.

Bonis, N.R., Kürschner, W.M. & J.H.A. van Konijnenburg - van Cittert. 2010. Changing CO2 conditions during the end-Triassic inferred from stomatal frequency analysis on Lepidopteris ottonis (Goeppert) Schimper and Ginkgoites taeniatus (Braun) Harris. Palaeogeography, Palaeoclimatalogy, Palaeoecology 295: 146-161.

Kustatscher,  E., Wachtler, M. & Van Konijnenburg-van Cittert, J.H.A. 2010. Lycophytes from the Middle Triassic (Anisian) locality Kühwiesenkopf (Monte prà della Vacca) in the Dolomites (Northern Italy). Palaeontology 53(3): 595-626.

van der Ham, Raymond W.J.M., Jagt, John W.M., Renkens. Sjir & Johanna H.A. van
Konijnenburg-van Cittert. 2010. Seed-cone scales from the upper Maastrichtian document the last occurrence in Europe of the Southern Hemisphere conifer family Araucariaceae. Palaeogeography, Palaeoclimatalogy, Palaeoecology 291 (3-4): 469-473.

Kustatscher,  E.,  Van Konijnenburg-van Cittert, J.H.A. & G. Roghi. 2010. Macrofloras and palynomorphs as possible proxies for palaeoclimatic and palaeoecological studies: A case study of the Fossilelagerstätte Kühwiesenkopf/Monte Pra della vacca (Olang Dolomites, N-Italy). Palaeogeography, Palaeoclimatalogy, Palaeoecology 291(1-2): 71-80.

Collinson, M.E., Barke, J., van der Burgh, J., van Konijnenburg-van Cittert, J. H. A., Heilmann-Clausen, C., Howard, L. E. & Brinkhuis, H. 2010. Did a single species of Eocene Azolla spread from the Arctic Basin to the southern North Sea?  Review of Palaeobotany and Palynology159:152–165.

Zijlstra, G., van Konijnenburg-van Cittert, J.H.A., Kunzmann, L., Bosma, H. & Jirí Kvacek. 2010. Proposal to conserve the name Geinitzia (fossil Coniferophyta). Taxon 59(1): 301-302.

Zavialova,  N., van Konijnenburg-van Cittert, J.H.A. and M. Zavada, 2009. The pollen exine ultrastructure of the bennettitalean bisexual flower Williamsoniella coronata.  International Journal of Plant Sciences 170(9): 1195–1200

M. Booi, I.M. van Waveren & J.H.A. van Konijnenburg-van Cittert, 2009.  The Jambi gigantopterids and their place in gigantopterid classification.  Bot. J. Lin. Soc. 161(3): 302-328.

Konijnenburg-van Cittert van, J.H.A., 2009. Fossiele planten sinds het Carboon. Gea 42 (3): 65-71.

M. Booi, I.M. van Waveren & J.H.A. van Konijnenburg-van Cittert, 2009. Comia and Rhachiphyllum from the Early Permian of Sumatra, Indonesia. Review of Palaeobotany and Palynology, 156: 418-435.

Zijlstra, G., Kustatscher, E. & J.H.A. van Konijnenburg-van Cittert, 2009. Proposal to conserve the name Sphenozamites (fossil Cycadophyta). Taxon  58 (3): 1016.

Kustatscher, E., Hemsley, A.R. & Van Konijnenburg-van Cittert, J.H.A., 2009. Lugardonia paradoxa gen. et sp. nov., a new strobilus from the Anisian flora of Kühwiesenkopf, the Dolomites,  Italy and its affinities with emphasis on spore ultrastructure. Review of Palaeobotany and Palynology, 156: 90-97.

Collinson,  M.E., Barke, J., van der Burgh, J. &  Johanna H. A. van Konijnenburg-van Cittert, 2009.  A new species of the freshwater fern Azolla (Azollaceae)  from the Eocene Arctic Ocean. Review of Palaeobotany and Palynology, 155 (1-2): 1-14.

Bosma, H.F., van Konijnenburg-van Cittert, J.H.A., van der Ham, R.W.J.M., van Amerom, H.W.J. & Hartkopf-Froeder, C., 2009. Conifers from the Santonian of Limburg, the Netherlands. Cretaceous Research  30(2): 483-495.

Schweitzer, H.-J., Schweitzer, U., Kirchner, M., Van Konijnenburg-Van Cittert, J.H.A., Van der Burgh, J. & Ashraf, R.A., 2009. The Rhaeto-Jurassic flora of Iran and Afghanistan. 14. Pterophyta – Leptosporangiatae. Palaeontographica Abt. B. 279, pts 1-6 p. 1-108, 50 plates

Kustatscher, E. & Van Konijnenburg-van Cittert, J.H.A., 2008. Lycophytes and horsetails from the Triassic Flora of Thale (Germany). Neues Jahrbuch für Geologie und Palaeontologie, Abhandlungen, 250(1): 65-77.

Van Waveren, I.M., Abbink, O., van Hoof, T. & van Konijnenburg-van Cittert, J.H.A., 2008. Revision of the Late Carboniferous palaeobotany of the De Lutte-06 well (Twente, the Netherlands), and its stratigraphical implications. Neth. J. of Geology/Geologie en Mijnbouw 87(4): 339-352.

M. Booi, I.M. van Waveren & J.H.A. van Konijnenburg-van Cittert, 2008. New material of Macralethopteris from the Jambi area of Middle Sumatra and its climatological implications. Review of Palaeobotany and Palynology, 152: 101-112.

Kustatscher, E. & Van Konijnenburg-van Cittert, J.H.A., 2008. Considerations on Phylladelphia strigata Bronn from the historica Raibl flora (Carnian, lower Upper Triassic, Italy). Geo.Alp 5: 69-81.

Wade-Murphy, J. & Van Konijnenburg-van Cittert, J.H.A. 2008. A revision of the Late Triassic Bintan flora from the Riau Archipelago (Indonesia). Scripta Geologica 136: 73-105.

van Konijnenburg-van Cittert, J.H.A., 2008. The Jurassic fossil plant record of the UK area. Proceedings of the Geologists' Association 119: 59-72.

Van Konijnenburg-van Cittert, J.H.A. 2007. Bernettiales van Konijnenburg-van Cittert, order nov.. In: Anderson, J.M., Anderson, H.M. & Cleal, C.J. (eds.) Brief history of gymnosperms: classification, biodiversity, phytogeography and ecology. Strelitzia 20: 208. 

Van Waveren, I. M. Iskandar, E., & van Konijnenburg-van Cittert, J.H.A., 2007. Composition and palaeogeography of  the Lower Permian Jambi flora from Sumatra. Scripta Geologica 135: 1-28.  

Christian Pott, Johanna H. A. van Konijnenburg-van Cittert, Hans Kerp, Michael Krings, 2007. Revision of the genus Pterophyllum (Cycadophyta: Bennettitales) in the Carnian (Late Triassic) flora from Lunz, Lower Austria. Review Palaeobotany and Palynology 147: 3-27 

Pott, C., Kerp, J.H.F., Zijlstra, G. & van Konijnenburg-van Cittert, J.H.A., 2007, Proposal to conserve the name Pterophyllum  (Foss., Bennettitales) with a conserved type. Taxon 56 (3): 966-967. 

Gea Zijlstra, Evelyn Kustatscher & Johanna H.A. van Konijnenburg-van Cittert. 2007,Proposals to conserve the name Schizoneura against Convallarites and S. Paradoxaagainst C. erecta (fossil Sphenopsida). Taxon 56(3): 965-966. 

Kustatscher,  E., Wachtler, M. & Van Konijnenburg-van Cittert, J.H.A., 2007. Horsetails and seedferns from the Anisian locality Kühwiesenkopf (Dolomites, Northern Italy). Palaeontology 50 (5): 1277-1298.

Van der Ham, R.W.J.M., van Konijnenburg-van Cittert, J.H.A., & Indeherberge, L. 2007. Seagrass foliage from the Maastrichtian type area (Late Cretaceous, Early Tertiary, NE Belgium, SE Netherlands). Review Palaeobotany and Palynology 144: 301-321.  

Kustatscher, E. & Van Konijnenburg-van Cittert, J.H.A., 2007. Taxonomical and Palaeogeographical considerations on the genus Ptilozamites with some comments onAnomozamites, Dicroidium, Pseudoctenis and Ctenozamites. Neues Jahrbuch für Geologie und Palaeontologie, Stuttgart, Bd. 243(1): 71-100.  

Tekleva, M., Krassilov, V.A. , Kvacek, J. & van Konijnenburg-van Cittert, J.H.A., 2006. Pollen Genus Eucommiidites: ultrastructure and affinities. Acta Palaeobotanica 46(2): 137-155.  

Roghi G., Kustatscher K., and van Konijnenburg-van Cittert J. H.A., 2006, Late Triassic Plants from the Julian Alps (Italy). Boll. Soc. Paleont. It., 45 (I); 133-140.  

Van Konijnenburg-van Cittert, J.H.A., Wachtler, M. & Kustatscher,  E., 2006. Pteridophytes from the Anisian locality Kühwiesenkopf (Dolomites, Northern Italy). Palaeontology 49(5): 943-968.

Kustatscher, E., Meller, B. & van Konijnenburg-van Cittert, J.H.A., 2006. Old treasures newly discovered: Scytophyllum  bergeri from the Ladinian of the Dolomites in the historical collections of the Geologische Bundesanstalt Wien. GeoAlp 3: 47-53. 

Kustatscher, E., Van Konijnenburg-van Cittert, J.H.A. & Gianolla, P. 2006. The Kühwiesenkopf/Monte Pra della Vacca (Prags/Braies Dolomites, Northern Italy): An attempt to reconstruct an Anisian (lower Middle Triassic) palaeoenvironment. 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota, 27-29.05.06, Manchester, Abstract and Proceedings Volume, p. 63-66, 164. 

Popa, M.E. & van Konijnenburg-van Cittert, J.H.A. 2006. Aspects of Romanian Early – Middle Jurassic palaeobotany and palynology. Part VII. Successions and floras. Progress in Natural Science, Volume 16 (Special Issue: Marine and non-marine Jurassic: correlation and boundary events): 203-212.  

Brinkhuis, H. Schouten, S., Collinson, M.E., Sluijs, A., Sinnighe Damsté, J.S., Dickens, G.R., Huber, M., Cronin, T.M., Onodera, J., Takahashi, K., Bujak, J.P., Stein, R., van der Burgh, J., Eldrett, J.S., Harding, I.C., Lotter, A.F., Sangiorgi, F., van Konijnenburg-van Cittert, J.H.A., de Leeuw, J.W., Matthiessen, J., Backman, J., Moran, K. & the Expedition 302 Scientists. 2006. Episodic fresh surface waters in the Eocene Arctic Ocean. Nature, 441: 606-609. 

Herngreen, G.F.W., van Konijnenburg-van Cittert, J.H.A., Oosterink, H.W. & van der Ham, R.W.J.M. 2005. Nieuwe geologische, palynologische en paleobotanische gegevens (Muschelkalk, Rhaetien-Lias en Oligoceen) uit de steengroeven van Winterswijk. Grondboor en Hamer 59 (4): 84-97. 

Herngreen, G.F.W., van Konijnenburg-van Cittert, J.H.A., &Oosterink, H.W. 2005. New geological data (Middle Triassic, Rhaetian-Liassic and Oligocene) of the Winterswijk quarry, the eastern Netherlands. Neth. J. of Geology/Geologie en Mijnbouw, 84 (4): 409-413.  

 Waveren, I.M.van, Hasibuan, F., Suyoko, Makmur, Boer, P.L.de, Chaney, D., Ueno, K., Booi, M., Iskandar, E.A.P., King, C., Leeuw, J.M.de & van Konijnenburg-van Cittert, J.H.A. 2005. Taphonomy, palaeobotany and sedimentology of the Mengkarang Formation (Early Permian, Jambi, Sumatra, Indonesia). In: Lucas, E. (ed.), The Nonmarine Permian. Bull. New Mexico Museum of Natural History and Science, 30: 333-341. 

DiMichele, W.A., van Konijnenburg-van Cittert, J.H.A., Looy, C.V. & Chaney, D. 2005.Equisetites from the Early Permian of North-Central Texas. In: Lucas, E. (ed.), The Nonmarine Permian. Bull. New Mexico Museum of Natural History & Science, 30: 56-59.  

Evelyn Kustatscher & Van Konijnenburg-van Cittert, Johanna H.A., 2005. The Ladinian Flora (Middle Triassic) of the Dolomites: palaeoenvironmental reconstructions and palaeoclimatic consideratieons. Geo. Alp. 2: 31-51. 

Looy, C.V., Collinson, M.E., Van Konijnenburg-van Cittert, J.H.A. & Visscher, H., 2005. The ultrastructure and botanical affinity of end-Permian spore tetrads. Int. J. Plant Sciences 166 (5): 875-887. 

Kustatscher, E., Wachtler, M. & Van Konijnenburg-van Cittert, J.H.A., 2004: Some additional and revised taxa from the Ladinian flora of the Dolomites,Northern Italy. Geo.Alp. 1: 57-70.

Van der Ham, R.W.J.M. & Van Konijnenburg-van Cittert, J.H.A., 2004. Coniferen uit de Campanien en Maastrichtien van Zuid-Limburg en omgeving. Natuurhistorisch Maandblad93: 26-32.

Konijnenburg- van Cittert, J. H. A. van, I. M. van Waveren & J. Jonkers, 2004. Catalogue of the holotypes in the collection of plant fossils in theNational Museum of Natural History, Leiden. Technical Bulletin Naturalis7.

Abbink, O.A., van Konijnenburg-van Cittert, J.H.A., & Visscher, H. 2004. A Sporomorph Ecogroup Model for the Northwest European Upper Jurassic Lower Cretaceous: Concepts and framework. Netherlands Journal of Geology83(1): 389-404.

Van der Ham, R.W.J.M., Van Konijnenburg-van Cittert, J.H.A. & Nieuwenhuis, E.A.P.M, 2004. Cunninghamites ubaghsii (Taxodiaceae?) from the Maastrichtian tyope area (Late Cretaceous, SE Netherlands) rediscovered. Bull. Inst. Royal Scienc. Natur. De Belgique, Sci. de la Terre74: 89-96.

Visscher,H. C. van Looy, M. E. Collinson, H. Brinkhuis, J. H.A. van Konijnenburg-van Cittert, W.M. Kürschner, & M.A. Sephton, 2004. Environmental Mutagenesis during the End-Permian Ecological Crisis. Proceedings National Academy of Sciences USA

Passoni, L. & J.H.A. van Konijnenburg-van Cittert, 2003. New taxa of fossil Carnian Plants from Mount Pora (Bergamasc Alps; Northern Italy). Rev. Palaeobot. Palynol. 123: 321-346.

Ham, R.W.J.M. van der, & J.H.A.van Konijnenburg-van Cittert, 2003. Rare conifers from the type area of the Maastrichtian (Upper Cretaceous, southeast Netherlands). Scripta Geologica 126: 111-119, 3figs., 2 pls.

Gee, C.T., Meyer, C. A. & J.H.A.van Konijnenburg-van Cittert, 2003. The first unequivocal sphenophytes from the Late Jurassic of northwestern Switzerland (late Oxfordian Vellerat Formation) - Palaeobotany and implications for facies interpretation. Courier Forschungs-Institut Senckenberg 241: 97-109.

Ham, R.W.J.M. van der, Konijnenburg-van Cittert, J.H.A. van, Burgh, J. van der & Herngreen, W., 2003. Brachyphyllum patens (Miquel) comb. nov.: remarkable conifer foliage from the Maastrichtian type area (Late Cretaceous, NE Belgium, SE Netherlands). Rev. Palaeobot. Palynol. 127: 77-97.

Collinson, M.E. & Van Konijnenburg-van Cittert, J.H.A (Eds.).2002 Ecology of Ferns through Time. Special Volume Rev. Palaeobot. Palynol. 119.

Van Konijnenburg-van Cittert, J.H.A., 2002. Ecology of some Jurassic ferns in Eurasia. Rev. Palaeobot. Palynol. 199: 113-124.

Konijnenburg-van Cittert, J.H.A. van, Schmeissner, S., S.& T. Hauptmann, 1998. Neue Ergebnisse zu Ctenozamites wolfiana (Pteridospermae) und Pseudoctenis prossii nov. spec. (Cycadophyta) aus dem Unteren Lias (Jura, Bayern). Documenta naturae 117: 13-33.

Barbacka, M. & J.H.A. van Konijnenburg-van Cittert, 1998. Sun and shade leaves in two Jurassic species of Pteridosperms. Rev. Palaeobot. Palynol.103: 209-211.

Guignard, G., Thévenard, F. & J.H.A. van Konijnenburg-van Cittert, 1998. Cuticle ultrastructure of the cheirolepidiaceous conifer Hirmeriella muensteri (Schenk)Jung. Rev. Palaeobot. Palynol.104: 115-141.

Looy, C.V., van Konijnenburg-van Cittert, J.H.A., Visscher, H. & Dilcher, D.L. 2001. Life in the end-Permian dead zone. PNAS (Proceedings National Academy of Sciences USA). 98 (14): 7879-7883

Van der Ham, R.W.J.M., Van Konijnenburg-van Cittert, J.H.A. & Van der Burgh, J. 2001. Taxodiaceous conifers from the Maastrichtian type area (Late Cretaceous, NE Belgium, SE Netherlands). Rev. Palaeobot. Palynol. 116: 233-250.

J.H.A.van Konijnenburg-van Cittert & Bandel, K.2001. Jurassic plants from Djebel Thi, Sinaï. Mitt. Geol.-Paläont. Inst. Univ. Hamburg 85: 47-64.

Van Konijnenburg-van Cittert, J.H.A., Schmeißner, S. & G. Dütsch. 2001. A newRhaphidopteris from the Lower Liassic of Bavaria, Germany. Acta Palaeobotanica 41(2): 109-115.

Van Konijnenburg-van Cittert, J.H.A., Miku V., & Pavsic, J. 2001. Pterophyllum(Cycadopsida) from Carnian beds in PoljaneValley. Geologija 44 (2): 317-323.

Van Waveren, I., J.H.A.van Konijnenburg-van Cittert, Van der Burgh, J. & Dilcher, D.L.: Macrofloral remains of the Leiva region (Colombia). Scripta geologica 123: 1-39.

J.H.A.van Konijnenburg-van Cittert & Bandel, K.2001. Jurassic plants from Djebel Thi, Sinaï. Mitt. Geol.-Paläont. Inst. Univ. Hamburg 85: 47-64.

Wachtler, M. & J.H.A.van Konijnenburg-van Cittert, 2000. The fossil Flora of the Wengen Formation (Ladinian) in the Dolomites (Italy). Beiträge zur Paläontologie, 25: 105-141.

Sarjeant, W.A.S., Musteikis, P., Abbink, O. & J.H.A. van Konijnenburg-van Cittert, 2000. A new record of Jurassic palynomorphs from Lithuania. Geologija 32: 76-87.

Schweitzer, H.-J., Kirchner, M. & J.H.A.van Konijnenburg-van Cittert, 2000. The Rhaeto-Jurassic flora of Iran and Asfghanistan. 12. Cycadophyta II. Nilssoniales. Palaeontographica Abt.B, 254: 1-63.

Manum, S, Van Konijnenburg-van Cittert, J.H.A. & Wilde, V., 2000: Tritaenia Mägdefrau et Rudolf, Mesozoic Sciadopitys-like leaves in mass accumulations. Rev. Palaeobot. Palynol., 109: 255-269.

Zijlstra, G. & van Konijnenburg-van Cittert, J.H.A., 2000 Proposal to conserve the name Araucarites C. Presl (Fossil Gymnospermae, Coniferales, Araucariaceae) against Araucarites Endl. (Fossil Gymnospermae, Coniferales). Taxon 49: 279-280.

J.H.A.van Konijnenburg-van Cittert, 2000: Parameters in leaf and cuticle morphology. The Palaeobotanist 49: 9-15.

J.H.A.van Konijnenburg-van Cittert, 2000. Osmundaceous spores throughout time. In: M.M.Harley, C.M.Morton and S.Blackmore (Eds.). Pollen and Spores: Morphology and Biology, pp. 435-449. Royal Botanic Gardens, Kew.

Popa, M.E. & Van Konijnenburg-van Cittert, J.H.A., 1999 (printed 2000). Aspects of Romanian Early Jurassic paleobotany and palynology. Part 1. In situ spores from the Getic Nappe, Banat, Romania. Acta Palaeobotanica, suppl.2: 181-193.

Thevenard, F., Guignard, G. & Van Konijnenburg-van Cittert, J.H.A., 1999 (printed 2000). Ultrastructural study of the cuticle of Himeriella muensteri. Acta Palaeobotanica, suppl.2: 233-235.

J.H.A.van Konijnenburg-van Cittert & H.S.Morgans, 1999. A guide to the Yorkshire Jurassic flora. Palaeontological Association. Field Guide to Fossils No.8; 134 pp.

J.H.A.van Konijnenburg-van Cittert & S. Schmeissner, 1999. Fossil insect eggs on Lower Jurassic plant remains from Bavaria (Germany). Palaeogeography, Palaeoclimatalogy, Palaeoecology 152: 215-223.

J.H.A.van Konijnenburg-van Cittert, 1999. On the evolution of fern spore architecture. In: M.H. Kurmann & A.R.Hemsley (eds.). The evolution of plant architecture; Whitstable Litho Printers Ltd., Whitstable, Kent, UK. pp. 279-287.

Wachtler, M. &  J.H.A.van Konijnenburg-van Cittert, 1998. Die Pflanzenwelt im Ladin. In: Stingl, V. & Wachtler, M. (eds.). Dolomiten, das Werden einer Landschaft. Naturmuseum Südtirol, Bozen.

Please contact the author when you are interested in one of the articles above and it is not available online or in the Naturalis repository.