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Dennis Uit de Weerd

Dr. D.R. (Dennis) Uit de Weerd, Research associate


Email: dennis.uitdeweerd@ou.nl
Phone: +31 (0)38 42 97 617
Room number: not applicable



Current positions

  • Assistent professor, School of Science, Open University Netherlands
  • Research associate,  Dept. Terrestrial Zoology, Naturalis Biodiversity Center
  • Research associate,  Laboratory for Molecular Systematics & Ecology, Academy of Natural Sciences Philadelphia


Research interest

Evolution and biogeography of land snails.




molecular phylogenetics, character evolution, biogeography

Current research topics

- Evolution and biogeography of the Caribbean land snail family Urocoptidae

- Evolution within the land snail family Clausilliidae




Available student projects


Uit de Weerd, D. R., Robinson, D. G., & Rosenberg. G. (2016). Evolutionary and biogeographical history of the land snail family Urocoptidae (Gastropoda: Pulmonata) across the Caribbean region. Journal of Biogeography 43 (4): 763–777.

Uit de Weerd, D. R., & Gittenberger, E. (2013). Phylogeny of the land snail family Clausiliidae (Gastropoda: Pulmonata). Molecular Phylogenetics and Evolution 67 (1): 201–216

Bertola, L. D., van Hooft, W. F., Vrieling, K., Uit de Weerd, D. R., York, D. S., Bauer, H., . . . de Iongh, H. H. (2011). Genetic diversity, evolutionary history and implications for conservation of the lion (Panthera leo) in West and Central Africa. Journal of Biogeography38: 1356–1367.

Uit de Weerd, D. R., & Gittenberger, E. (2010). Molecular phylogeny of the land snail family Clausiliidae Tropical Natural History, supplement 3: 176.

Uit de Weerd, D.R., Schneider, D. & E. Gittenberger, E. (2009). Molecular phylogenetic relationships of Inchoatia taxa. Zoologische Mededelingen83: 589-592 

Gittenberger, E. & Uit de Weerd, D.R. (2009). Summarizing data on the Inchoatia taxa,including Inchoatia megdova bruggeni subspec. nov. (Gastropoda, Pulmonata, Clausiliidae).Zoologische Mededelingen 83: 577-587. 

Uit de Weerd, D.R. (2008). Delimitation and phylogenetics of the highly diverse land snail family Urocoptidae (Gastropoda, Pulmonata) based on 28S rRNA sequence data: A reunion with Cerion. Journal of Molluscan Studies 74: 317-329.

Uit de Weerd, D.R. (2007). Phylogeography and evolution of the Caribbean land snail family Urocoptidae (Gastropoda, Pulmonata): radiation in isolation. Abstracts World Congress of Malacology Antwerp 2007 

Gittenberger, E & Uit de Weerd, D.R. (2006). Reconsidering the generic position of the species once classified in CarinigeraIsabellaria and Sericata (Gastropoda, Pulmonata, Clausiliidae, Alopiinae). Basteria 70: 57-66. 

Uit de Weerd, D.R. , Groenenberg, D.S.J., Schilthuizen, M. & Gittenberger, E. (2006).Reproductive character displacement by inversion of  coiling in clausiliid snails (Gastropoda, Pulmonata). Biological Journal of the Linnean Society 88: 155-164. 

Gittenberger, E. & Uit de Weerd, D.R. (2006). The genus Inchoatia validated and three mistakes corrected.  Basteria 70: 131-132. 

Uit de Weerd, D.R, Schneider, D. & Gittenberger, E. (2005). The provenance of the Greek land snail species Isabellaria pharsalica: molecular evidence of recent long-distance dispersal.Journal of Biogeography 32: 1571-1581

Gittenberger, E & Uit de Weerd, D.R. (2005) Babylonia and Zemiropsis (Gastropoda, Caenogastropoda, Babyloniidae), anatomy, shell morphology, distribution and DNA. Journal of Conchology 38 ( 6): 649-652  

Uit de Weerd, D.R. & Gittenberger, E. (2005) Towards a monophyletic genus Albinaria(Gastropoda, Pulmonata): the first molecular study into the phylogenetic position of easternAlbinaria species. Zoological Journal of the Linnean Society 143 (4): 531-542. 

Uit de Weerd, D.R., Piel, W.H. & Gittenberger, E. (2004) Widespread polyphyly among Alopiinae snail genera: when phylogeny mirrors biogeography more closely than morphology.Molecular Phylogenetics and Evolution 33: 533-548. 

Uit de Weerd, D.R. (2004) Molecular phylogenetic history of eastern Mediterranean Alopiinae, a group of morphologically indeterminate land snails. PhD Thesis, Leiden University, The Netherlands. pp 1-119. (go to Leiden University Digital Repository)

Uit de Weerd, D.R., Gittenberger, E. (2004) Re-evaluating Carinigera: Molecular data overturn the current classification within the Clausiliid subfamily Alopiiinae (Gastropoda, Pulmonata).Journal of Molluscan Studies 70 (4): 305-318 

Gittenberger, E & Uit de Weerd, D.R. (2004) Polytypic Carinigera buresi in NE Greece (Gastropoda, Pulmonata, Clausiliidae). Basteria 67: 115-126. 

Caspers, G.-J., Uit de Weerd, D., Wattel, J. & de Jong, W.W. (1997). Alfa-crystallin sequences support a Galliform/Anseriform clade. Molecular Phylogenetics and Evolution 7: 185-188