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Annick S. Lang

A.S. (Annick) Lang, MSc., PhD Student - Botany, Bryology


Email: annick.lang@naturalis.nl
Phone: +31 (0)71-52 74 728
Room number: 1.01, Nieuwenhuizenweg 19
“Go down on your knees, put your nose in a moss cushion, and you'll discover the exiting world of bryophytes!”


Research interest

Bryophytes are small organisms that are found in every ecosystems except oceans. Their haplo-diplont life cycle and their ancestral phylogenetic position make them ideal subject for evolution, phylogeographic, biodiversity and population genetic studies.


bryophyte, taxonomy, phylogeny, systematics

Current research topics

Dicranum is a common and widespread moss genus in the Northern hemisphere. Although the species of this genus can be considered well-defined, many species are morphologically highly polymorphicand hence led taxonomical confusions.

The aims of this PhD project are to understand the relationship of closely related Dicranum species using molecular methods. I am furthermore interested in clarifying species circumscription, especially within species complexes,using barcoding and automated species delimitation methods.

The promotor of my PhD is Prof. Dr. Erik Smets. The co-promotor of my PhD is Dr. Michael Stech.


Not applicable.



Not applicable.

2012 Daily supervisor. L. E. van Dijk. Gender frequencies in populations of Dicranum scoparium.

Available student projects


Naturalis Repository

Lang A. S., D. Tubanova & M. Stech(in press), Species delimitations in the Dicranum acutifolium complex (Dicranaceae, Bryophyta), Journal of Bryology 

Lang A. S.,  Kruijer J. D. & M. Stech. (2014), DNA barcoding of arctic bryophytes – an example from the moss genus Dicranum (Dicranaceae, Bryophyta), Polar Biology, doi: 10.1007/s00300-014-1509-7 (2.006)

Lang A. S. & M. Stech (2014) What's in a name? Disentangling the Dicranum scoparium species complex (Dicranaceae, Bryophyta), Systematic Botany 39:369-379 

Price M. J.& A. S. Lang, 2011, Dicranum tauricum Sapjegin ( Dicranaceae ) new for the Canton of Geneva, Switzerland, Meylania 47: 29-33

Lang, A & Y. Naciri, 2010. New chloroplast primers for intraspecific variation in Dicranum scoparium Hedw. (Dicranaceae) and amplification success in other bryophyte species. Molecular Ecology Resources, 10: 735-737