The Terrestrial Gastropods of Sabah (North Borneo)

Taxonomic disclaimer: This web page is an informal publication and is not valid for taxonomic or nomenclatural purposes under the International Code of Zoological Nomenclature.

Below follows a list, with some illustrations, of all species of terrestrial snails and slugs currently known from the Malaysian state of Sabah. A monograph is in preparation, in which a large number of new species will be described. Those species are below referred to with "Vermeulen, Schilthuizen & Liew MS". Voucher specimens of most of the species listed here are found in the BORNEENSIS collection of the Institute for Tropical Biology and Conservation and in the private collection of Jaap Vermeulen, Leiden, the Netherlands.

Species list

Assimineidae

Acmella cyrtoglyphe Vermeulen, Schilthuizen & Liew MS
Acmella compressa Vermeulen, Schilthuizen & Liew MS
Acmella minutissima Maassen 200?
Acmella nana Vermeulen, Schilthuizen & Liew MS
Acmella ovoidea Vermeulen, Schilthuizen & Liew MS
Acmella polita Von Moellendorff 1887
Acmella striata Vermeulen, Schilthuizen & Liew MS
Acmella subcancellata Vermeulen, Schilthuizen & Liew MS
Acmella umbilicata Vermeulen, Schilthuizen & Liew MS
Anaglyphula pellislacertae Vermeulen, Schilthuizen & Liew MS
Assiminea brevicula (Pfeiffer 1854)
Assiminea nitida (Pease 1864)
"Cyclotropis ""V12713""" Vermeulen, Schilthuizen & Liew MS
Cyclotropis bedaliensis (Rensch 1934)

Cyclophoridae

Alycaeus globosus H. Adams 1870
Alycaeus jagori Von Martens 1859
Chamalycaeus everetti (Godwin Austen 1889)
Chamalycaeus specus (Godwin Austen 1889)
"Chamalycaeus ""V1599""" Vermeulen, Schilthuizen & Liew MS
Cyclophorus kinabaluensis E.A. Smith 1895
Cyclophorus perdix borneensis (Metcalfe 1851)
Ditropopsis constricta Vermeulen, Schilthuizen & Liew MS
Ditropopsis davisoni Vermeulen, Schilthuizen & Liew MS
Ditropopsis imadatei (Habe 1965)
Ditropopsis koperbergi (Zilch 1955)
Ditropopsis trachychilus Vermeulen, Schilthuizen & Liew MS
Ditropopsis tyloacron Vermeulen, Schilthuizen & Liew MS
Japonia alticola (Laidlaw 1937)
Japonia amauroconus Vermeulen, Schilthuizen & Liew MS
Japonia anceps Vermeulen, Schilthuizen & Liew MS
Japonia balabacensis balabacensis (E.A. Smith 1895)
Japonia balabacensis adscendens Vermeulen, Schilthuizen & Liew MS
Japonia compressa (Von Moellendorff 1897)
Japonia dido Godwin Austen 1889
Japonia inornatus (E.A. Smith 1893)
Japonia janus Vermeulen, Schilthuizen & Liew MS
Japonia jucunda (E.A. Smith 1893)
Japonia keppeli (Godwin Austen 1889)
Japonia kinabaluensis (E.A. Smith 1895)
Japonia lissocoryphe Vermeulen, Schilthuizen & Liew MS
Japonia minutior Vermeulen, Schilthuizen & Liew MS
Japonia proprium (Fulton 1905)
Japonia smithi (Kobelt & Von Moellendorff 1897)
Japonia subrudis Vermeulen, Schilthuizen & Liew MS
Japonia taphroconus Vermeulen, Schilthuizen & Liew MS
Japonia trilirata (Pfeiffer 1852)
Japonia whiteheadi (E.A. Smith 1887)
Leptopoma undatum (Metcalfe1851)
Leptopoma pellucidum (Grateloup 1840)
Leptopoma sericatum (Pfeiffer 1851)
Leptopoma atricapillum (Sowerby 1843)
Opisthoporus birostris (Pfeiffer 1854)
Opisthoporus iris (Godwin Austen 1889)
"Opisthoporus ""V7578""" Vermeulen, Schilthuizen & Liew MS
Platyraphe bongaoensis (E.A. Smith 1894)
Platyraphe bicolor (Von Martens 1903)
Pterocyclos amabilis Fulton 1905
Pterocyclos tenuilabiatus (Metcalfe 1851)
Pterocyclos trusanensis (Godwin Austen 1889)

Diplommatinidae

Arinia biplicata Vermeulen 1996
Arinia boreoborneensis Vermeulen 1996
Arinia borneensis E.A. Smith 1893
Arinia brevispira orientalis Vermeulen 1996
Arinia brevispira brevispira Vermeulen 1996
Arinia clausa Vermeulen 1996
Arinia cylindrica cylindrica Vermeulen 1996
Arinia cylindrica crassilabris Vermeulen 1996
Arinia dentifera Vermeulen 1996
Arinia oviformis Vermeulen 1996
Arinia paricostata Vermeulen 1996
Arinia pertusa Vermeulen 1996
Arinia simplex Vermeulen 1996
Arinia stenotrochus pachystoma Vermeulen 1996
Arinia stenotrochus strenotrochus Vermeulen 1996
Arinia streptaxiformis Vermeulen 1996
Arinia turgida Vermeulen 1996
Diplommatina adversa (H. & A. Adams 1851)
Diplommatina antheae Vermeulen 1993
Diplommatina asynaimos Vermeulen 1993
Diplommatina cacuminulus Vermeulen 1993
Diplommatina calvula Vermeulen 1993
Diplommatina centralis Vermeulen 1993
Diplommatina cyclostoma Von Moellendorff 1897
Diplommatina cyrtorhitis Vermeulen 1993
Diplommatina electa Fulton 1895
Diplommatina gomantongensis E.A. Smith 1894
Diplommatina isseli Godwin Austen 1889
Diplommatina madaiensis Vermeulen 1993
Diplommatina oedogaster Vermeulen 1993
Diplommatina plecta Fulton 1901
Diplommatina recta E.A. Smith 1895
Diplommatina rubicunda (Von Martens 1864)
Diplommatina rubra Godwin Austen 1889
Diplommatina soror Vermeulen 1993
Diplommatina strongyla Vermeulen 1993
Diplommatina sykesi Fulton 1901
Diplommatina tenuilabiata Fulton 1901
Diplommatina toretos Vermeulen 1993
"Diplommatina ""V9928""" Vermeulen, Schilthuizen & Liew MS
"Diplommatina ""Trusmadi""" Vermeulen, Schilthuizen & Liew MS
Diplommatina whiteheadi E.A. Smith 1898
Opisthostoma bihamulatum Vermeulen 1994
Opisthostoma brachyacrum lambii Vermeulen 1991
Opisthostoma brevituba Vermeulen 1994
Opisthostoma concinnum Fulton 1901
Opisthostoma cyrtopleuron Vermeulen 1994
Opisthostoma decrespignyi (H. Adams 1865)
Opisthostoma dormani Vermeulen 1994
Opisthostoma fraternum E.A. Smith 1905
Opisthostoma hailei Solem 1964
Opisthostoma inornatum Vermeulen 1994
Opisthostoma javanicum Van Benthem Jutting 1932
Opisthostoma jucundum E.A. Smith 1893
Opisthostoma lechria Vermeulen 1991
Opisthostoma lissopleuron bigibbum Vermeulen 1994
Opisthostoma lissopleuron lissopleuron Vermeulen 1994
Opisthostoma mirabile E.A. Smith 1893
Opisthostoma obliquedentatum Vermeulen 1994
Opisthostoma otostoma Boettger 1893
Opisthostoma perspectivum Vermeulen 1994
Opisthostoma simplex Fulton 1901
Opisthostoma telestoma Vermeulen 1991
"Opisthostoma ""V7991""" Vermeulen, Schilthuizen & Liew MS
"Opisthostoma ""V7691""" Vermeulen, Schilthuizen & Liew MS

Helicinidae

Aphanoconia dichroa (Von Moellendorff 1890)
Aphanoconia usukanensis (Godwin Austen 1889)
Geophorus agglutinans (Sowerby 1842)
Sulfurina euchromia Wagner 1905
Sulfurina martensi (Issel 1874)

Hydrocenidae

Georissa bangueyensis E.A. Smith 1895
Georissa borneensis E.A. Smith 1895
Georissa everetti E.A. Smith 1895
Georissa filiasaulae Haase & Schilthuizen 2006
Georissa gomantongensis E.A. Smith 1893
Georissa scalinella (Van Benthem Jutting) 1966
Georissa saulae (Van Benthem Jutting) 1966
Georissa similis E.A. Smith 1893
"Georissa ""V1829""" Vermeulen, Schilthuizen & Liew MS
"Georissa ""V7623""" Vermeulen, Schilthuizen & Liew MS
"Georissa ""V8081""" Vermeulen, Schilthuizen & Liew MS
Georissa williamsi Godwin Austen 1889

Pupinidae

Pupina hosei Godwin Austen 1889

Truncatellidae

Truncatella guerini A. & J.B. Villa 1841

Veronicellidae

Laevicaulis alte (Ferussac) 1821
Sarasinula plebeia (Fischer) 1868
Valiguna flava (Heynemann) 1885

Achatinellidae

Elasmias globulosum Quadras & Moellendorff ex Zilch 1962
Elasmias manilense (Dohrn) 1863
Tornatellinops moluccana (Boettger) 1891

Achatinidae

Achatina fulica Bowdich 1822

Ariophantidae

Durgella densestriata Blanford 1863
Dyakia hugonis (Pfeiffer) 1863
Everettia jucunda (Pfeiffer 1863)
Everettia subconsul (Smith 1887)
Everettia themis (Smith 1895)
Everettia klemmantanica Gude 1918
Everettia corrugata corrugata Laidlaw 1937
Everettia corrugata williamsi Liew, Schilthuizen & Vermeulen 2009
Everettia layanglayang Liew, Schilthuizen & Vermeulen 2009
Everettia lapidini Liew, Schilthuizen & Vermeulen 2009
Everettia interior Liew, Schilthuizen & Vermeulen 2009
Everettia paulbasintali Liew, Schilthuizen & Vermeulen 2009
Everettia jucundior Liew, Schilthuizen & Vermeulen 2009
Everettia jasilini Liew, Schilthuizen & Vermeulen 2009
Everettia safriei Liew, Schilthuizen & Vermeulen 2009
Everettia occidentalis Liew, Schilthuizen & Vermeulen 2009
Everettia planispira Liew, Schilthuizen & Vermeulen 2009
Everettia monticola Liew, Schilthuizen & Vermeulen 2009
Everettia dominiki Liew, Schilthuizen & Vermeulen 2009
Hemiplecta humphreysiana (Godwin Austen) 1891
Hemiplecta egeria E.A. Smith 1895
Ibycus rachelae Schilthuizen & Liew 2008
Kalamantania whiteheadi (Godwin Austen) 1891
Macrochlamys indica
Macrochlamys tersa
Microcystina callifera Vermeulen, Schilthuizen & Liew MS
Microcystina consobrina Van Benthem Jutting 1959
Microcystina gratilla Van Benthem Jutting 1950
Microcystina lissa Vermeulen, Schilthuizen & Liew MS
Microcystina microrhynchus Vermeulen, Schilthuizen & Liew MS
Microcystina muscorum Van Benthem Jutting 1959
Microcystina physotrochus Vermeulen, Schilthuizen & Liew MS
Microcystina sinica Von Moellendorff 1885
Microcystina striatula Vermeulen, Schilthuizen & Liew MS
Microcystina subplanata Vermeulen, Schilthuizen & Liew MS
Microparmarion pollonerai Collinge & Godwin Austen 1895
Microparmarion simrothi Collinge & Godwin Austen 1895
Parmarion martensi Simroth 1893
Parmarion n. sp.
Philippinella moellendorffi Collinge 1899
Rhinocochlis moluensis
Sabalimax pantherina Tillier & Bouchet 1988
Sabalimax cyanantyx Tillier & Bouchet 1988
Vitrinodyakia viridis Vermeulen, Schilthuizen & Liew MS
Vitrinula decrespignyi (Higgins) 1868
"Vitrinula ""V7460""" Vermeulen, Schilthuizen & Liew MS

Bradybaenidae

Camaenidae

Amphidromus adamsi adamsi (Reeve) 1848
Amphidromus adamsi pictus Fulton 1896
Amphidromus hamatus Fulton 1896
Amphidromus martensi Boettger 1894
Chloritis dulcissima (E.A. Smith) 1898
Chloritis kinibalensis (Kobelt) 1894
Chloritis plena (Godwin Austen) 1891
Chloritis sibutuensis (E.A. Smith) 1894
Chloritis tomentosa (Pfeiffer in Reeve) 1854
Ganesella acris (Benson) 1859
Ganesella tigaensis (Godwin Austen) 1891
Landouria rotatoria (Von dem Busch) 1842
Obba marginata (Mueller) 1774
Planispira aff. loxotropis
Trachia pudica (Godwin Austen) 1891

Carychiidae

Carychium javanum Von Moellendorff 1897
Coilostele inquirenda Rensch 1935

Charopidae

Beilania philippinensis (Semper) 1874
Pilsbrycharopa kobelti (Boettger) 1908
Discocharopa aperta (Von Moellendorff) 1888
Charopa argos Vermeulen, Schilthuizen & Liew MS
Charopa infrastriata Vermeulen, Schilthuizen & Liew MS
Charopa jugalis Vermeulen, Schilthuizen & Liew MS
Charopa lissobasis Vermeulen, Schilthuizen & Liew MS
Charopa nubigena Vermeulen, Schilthuizen & Liew MS
Charopa platycephala Vermeulen, Schilthuizen & Liew MS
Charopa turgidula Vermeulen, Schilthuizen & Liew MS

Clausiliidae

Phaedusa filicostata filialis (Von Martens) 1903

Endodontidae

Philalanka anomphala Vermeulen, Schilthuizen & Liew MS
Philalanka brunnescens Vermeulen, Schilthuizen & Liew MS
Philalanka kusana (Aldrich) 1889
Philalanka moluensis (E.A. Smith) 1893
Philalanka obscura Vermeulen, Schilthuizen & Liew MS
Philalanka rugulosa Vermeulen, Schilthuizen & Liew MS
Philalanka thienemanni Rensch 1932

Euconulidae

Kaliella accepta (E.A. Smith) 1895
Kaliella barrakporensis (Pfeiffer) 1852
Kaliella calculosa (Gould) 1852
Kaliella dendrobates (Tillier & Bouchet) 1988
Kaliella dendrophila (Van Benthem Jutting) 1950
Kaliella doliolum (Pfeiffer) 1846
Kaliella gregaria (Tillier & Bouchet) 1988
Kaliella humilis (Tillier & Bouchet) 1988
Kaliella kinabaluensis (Tillier & Bouchet) 1988
Kaliella microconus (Mousson) 1865
Kaliella microeceos Vermeulen, Schilthuizen & Liew MS
Kaliella nephelophila (Tillier & Bouchet) 1988
Kaliella punctata Vermeulen, Schilthuizen & Liew MS
Kaliella scandens (Cox) 1872
Kaliella subplicata Vermeulen, Schilthuizen & Liew MS
Queridomus conulus (Blanford) 1865
"Rahula ""V9667""" Vermeulen, Schilthuizen & Liew MS

Ferussaciidae

Cecilioides caledonica (Crosse) 1867

Helicodiscidae

Stenopylis coarctata (Von Moellendorff) 1894

Limacidae

Deroceras laeve (Mueller) 1774

Rathouissidae

Atopos sp. 1
Atopos n. sp.
Atopos punctata Collinge 1902

Philomycidae

Meghimatium striatum (Van Hasselt 1824)
Meghimatium uniforme Laidlaw 1937
Meghimatium pictum (Stolyczka 1872)

Punctidae

Paralaoma albella Vermeulen, Schilthuizen & Liew
Paralaoma angusta Vermeulen, Schilthuizen & Liew
Paralaoma gracilitesta (Van Benthem Jutting) 1959
Paralaoma servilis (Shuttleworth) 1852

Rhytididae

"Macrocycloides ""V2877""" Vermeulen, Schilthuizen & Liew MS

Streptaxidae

Diaphera helenae Vermeulen 1990
Diaphera wilfordii ectyphus Vermeulen 1990
Diaphera wilfordii wilfordii Dance 1970
Huttonella bicolor (Hutton) 1834

Subulinidae

Allopeas clavulinum (Potiez & Michaud) 1838
Allopeas gracile (Hutton) 1834
"Borneopeas ""V1222""" Vermeulen, Schilthuizen & Liew MS
Opeas hannense (Rang) 1831
Paropeas achatinaceum (Pfeiffer) 1846
Subulina octona (Bruguiere) 1789

Trochomorphidae

Bertia brookei (Adams & Reeve) 1848
Geotrochus bongaoensis (E.A. Smith) 1894
Geotrochus conicoides (Metcalfe 1851) 1851
Geotrochus kinabaluensis
Geotrochus kitteli Vermeulen, Schilthuizen & Liew, MS
Geotrochus labuanensis (Pfeiffer 1863)
Geotrochus meristotrochus Vermeulen, Schilthuizen & Liew MS
Geotrochus obscura Vermeulen, Schilthuizen & Liew MS
Geotrochus pallida (E.A. Smith 1895)
Geotrochus rhysa Tillier & Bouchet 1988
"Geotrochus ""Tawau"""
Geotrochus whiteheadi (E.A. Smith 1895)
Videna bicolor (Von Martens 1864)
Videna froggatti (Iredale 1941)
Videna metcalfei (Pfeiffer 1845)
Videna repanda (Von Moellendorff 1890)
Videna timorensis (Von Martens 1867)

Vertiginidae

Boysidia salpinx Thompson & Dance 1983
Gastrocopta avanica (Benson) 1863
Gastrocopta pediculus
Gastrocopta recondita (Tapparone-Canefri) 1883
Gastrocopta servilis (Gould) 1843
Nesopupa malayana (Issel) 1874
Nesopupa moreleti (Brown) 1870
Ptychopatula circumlitum (Hedley) 1897
Ptychopatula orcella (Stoliczka) 1873
Ptychopatula orcula (Benson) 1850
Pupisoma pulvisculum (Issel) 1874
Pupisoma unidentata (Godwin Austen) 1910

Zonitidae

Zonitoides arboreus (Say) 1819

Saturday, October 3, 2009